Power Botanizing the Tallgrass Prairie in the Northern Heartland

For the Love of Grass VI

The sixth botanizing trek for the “Love of Grass” enjoyed a special rendezvous location, assembling at the home of trek participant Robert MacLauchlan in Lincoln, Nebraska.  By five o’clock in the afternoon on August 17, 2013 the six members from the 2012 trek were enjoying fellowship and a cookout provided by Bob’s daughter and son-in-law Nancy and Andy Bess. Bob is blessed with children close by.

The sixth trek was originally scheduled for a spring sojourn through the foothills of California. Scheduling difficulties led to this later agenda, put together by Jack Carlson. He and plant identifying wizard Larry Holzworth constituted the botanizing power, with Wendall Oaks and Wendell Hassell in major support roles, and added photographic responsibilities. Curtis Sharp and Bob rounded out the team, supplying wisdom and good looks, of questionable botanizing value but useful at the Prairie Chick in Okobogi, Iowa. 

Our inventory mission was the same as previous treks, at each site to identify every plant within a ‘reasonable’ area, consisting of one to several acres, and leave only when satisfied that this had been accomplished. Increasing skill levels, shorter distances between sites and motel reservations via cell phones resulted in slightly shorter workdays, much to the delight of the ancient ones, 83 and 85.  

During the seven days of inventorying the team traveled 1,534 miles, inventoried 17 sites at 13 locations, and identified as many as 127 species at one site, correctly or incorrectly identified 1,083 plants of 312 species, saw three plus million acres of corn and soybeans, scheduled the 2014 trek, and four of the trekkers endured an epidemic of chigger bites.  At least we had evidence for our spouses and significant others of the fun we had. Of the species we identified 39% occurred only once, 15% twice, 9% three times and 4 through 16 times 37%. The top 20 species occurring at the 17 inventory sites were:

Scientific NameFrequency
Bromus inermis Leyss (smooth brome)16
Andropogon gerardii Vitman (big bluestem)15
Amorpha canescens Pursh (leadplant)14
Ascelpias syriaca L. (common milkweed)14
Solidago altissima L. (Canada goldenrod)14
Ambrosia psilostachya DC. (Cuman ragweed)13
Artemisia ludoviciana Nutt. (white sagebrush)13
Poa pratensis L. (Kentucky bluegrass)13
Rosa arkansana Porter (prairie rose)13
Schizachyrium scoparium (Michx.) Nash (little bluestem)13
Echinacea angustifolia DC. (purple coneflower)12
Oligoneuron rigidum (L.) Small (stiff goldenrod)12
Panicum virgatum L. (switchgrass)12
Bouteloua curtipendula (Michx.) Torr. (sideoats grama)11
Dalea purpurea Vent. (purple prairieclover)11
Liatris aspera Michx. (tall blazing star)11
Spartina pectinata Bosc ex Link (prairie cordgrass)11
Symphoricarpos occidentalis Hook. (western snowberry)11
Artemisia campestris L. (field sagebrush)10
Melilotus officinalis (L.) Lam. (yellow sweetclover)10

Species and common names were taken from PLANTS (http://plants.usda.gov).  Most photographs were taken at the true site although the same species appeared at multiple sites. 

Some members of the trek team were struck by the notion of having enjoyed their plant materials profession so much that they would tramp through endless prairies just is see what plants were there, concluding they were greatly blessed. The trek schedule of years to come supports this point of view.  Proposed rendezvous for 2014 is Sacramento, California, on March 16, 2014. The objective is to find remnant perennial grasses in the annual grasslands surrounding the Central Valley.

During the cookout, Curtis Sharp provided the team with a pre-inventory he took at the hotel while waiting for the rest of the team to arrive that day.  Hence the following:

Inventory Site 0 – Horizon Inn, Lancaster County, NE

Scientific NameCommon NameTypeOrigin
Apocynum cannabinum L.IndianhempForbNative
Ascelpias incarnata L.swamp milkweedForbNative
Bromus inermis Leyss.smooth bromeGraminoidIntroduced
Cicuta maculata L.spotted water hemlockForbNative
Convolvulus arvensis L.field bindweedForbIntroduced
Gaura parviflora Douglas ex Lehm.velvetweedForbNative
Morus alba L.white mulberryTreeIntroduced
Physalis virginiana Mill.Virginia groundcherryForbNative
Vernonia fasciculata Michx.prairie ironweedForbNative

Day 1 – Sunday, August 18, 1913

By 8:30 AM bags were loaded, Bob Mac had been picked up and designated driver Oaks and navigator Carlson, with iPad in hand, headed for our first inventory site a short jaunt southwest of Lincoln. The air was clear, the traffic nil, the anticipation high.  

Inventory Site 1 – Spring Creek Prairie and Audubon Center, Lancaster County, NE

Spring Creek Prairie Audubon Center is an 808-acre tallgrass prairie nature preserve located 20 minutes southwest of Lincoln. Nearly 650 acres have never been plowed. This area contains tallgrass prairie, ponds and wetlands, wildflowers and grasses, a diverse assortment of birds and other wildlife, peaceful surroundings, and beautiful scenic vistas. This was a perfect place to start our trek.

Scientific NameCommon NameTypeOrigin
Abutilon theophrasti MedikvelvetleafForbIntroduced
Ambrosia artemisiifolia L.annual ragweedForbNative
Ambrosia psilostachya DC.Cuman ragweedForbNative
Ambrosia trifida L.great ragweedForbNative
Amorpha canescens PurshleadplantLegumeNative
Amorpha fruticosa L.false indigo bushShrubNative
Andropogon gerardii Vitmanbig bluestemGraminoidNative
Artemisia campestris L.field sagewortForbNative
Artemisia ludoviciana Nutt.white sagebrushForbNative
Ascelpias incarnata L.swamp milkweedForbNative
Ascelpias syriaca L.common milkweedForbNative
Asclepias tuberosa L.butterfly milkweedForbNative
Asclepias verticillata L.whorled milkweedForb Native
Bouteloua curtipendula (Michx.) Torr.sideoats gramaGraminoidNative
Bouteloua dactyloides (Nutt.) J.T. ColumbusbuffalograssGraminoidNative
Bouteloua gracilis (Willd. ex Kunth) Lag. ex Griffithsblue gramaGraminoidNative
Brickellia eupatorioides (L.) Shinnersfalse bonesetForbNative
Bromus arvensis L.field bromeGraminoidIntroduced
Bromus inermis Leyss.smooth bromeGraminoidIntroduced
Callirhoe involucrata (Torr. & A. Gray) A. Graypurple poppymallowForbNative
Celtis occidentaliseastern hackberryTreeNative
Chamaecrista fasciculata (Michx.) Greenepartridge peaLegumeNative
Chamaesyce nutans (Lag.) SmalleyebaneForbNative
Chenopodium standleyanum AellenStandley’s goosefootForbNative
Cirsium flodmanii (Rydb.) ArthurFlodman’s thistleForbNative
Cirsium undulatum (Nutt.) Spreng.wavyleaf thistleForbNative
Conium maculatum L.poison hemlockForbIntroduced
Convolvulus arvensis L.field bindweedForbIntroduced
Conyza canadensis (L.) CronquistCanadian horseweedForbNative
Cornus drummondii C.A. Mey.roughleaf dogwoodShrubNative
Cynodon dactylon (L.) Pers.bermudagrassGraminoidIntroduced
Dalea candida Michx. ex Willd.white prairiecloverLegumeNative
Dalea purpurea Vent.purple prairiecloverLegumeNative
Datura stramonium L.jimsonweedForbIntroduced
Daucus carota L.Queen Anne’s laceForbIntroduced
Desmodium illinoense A. GrayIllinois ticktrefoilLegumeNative
Digitaria sanguinalis (L.) Scop.hairy crabgrassGraminoidIntroduced
Echinacea angustifolia DC.blacksamson echinaceaForbNative
Elymus canadensis L.Canada wildryeGraminoidNative
Elymus repens (L.) GouldquackgrassGraminoidIntroduced
Equisetum laevigatum A. Braunsmooth horsestailForbNative
Eragrostis spectabilis (Pursh) Steud.purple lovegrassGraminoidNative
Erigeron strigosus Muhl. ex Willd.prairie fleabaneForbNative
Euphorbia dentata Michx.toothed spurgeForbNative
Euphorbia marginata Purshsnow on the mountainForbNative
Fraxinus pennsylvanica Marshallgreen ashTreeNative
Gaura longiflora Spachlongflower beeblossomForbNative
Gaura parviflora Douglas ex Lehm.velvetweedForbNative
Gleditsia triacanthos L.honeylocustTreeNative
Glycyrrhiza lepidota PurshAmerican licoriceLegumeNative
Grindelia squarrosa (Pursh) Dunalcurlycup gumweedForbNative
Helianthus annuus L.common sunflowerForbNative
Helianthus grosseserratus M. Martenssawtooth sunflowerForbNative
Helianthus maximiliani Schrad.Maximillian sunflowerForbNative
Helianthus pauciflorus Nutt. ssp. pauciflorus stff sunflowerForbNative
Heliopsis helianthoides (L.) Sweetsmooth oxeyeForbNative
Juniperus virginiana L.eastern redcedarTreeNative
Koeleria macrantha (Ledeb.) Schult.prairie junegrassGraminoidNative
Lactuca serriola L.prickly lettuceForbIntroduced
Lespedeza capitata Michx.roundhead lespedezaLegumeNative
Liatris punctata Hook.dotted blazing starForbNative
Liatris pycnostachya Michx.prairie blazing starForbNative
Linum sulcatum Riddellgrooved flaxForbNative
Lotus unifoliolatus (Hook.) Benth. var. unifoliolatusAmerican bird’s-foot trefoilLegumeNative
Medicago sativa L. alfalfaLegumeIntroduced
Monarda fistulosa L.wild bergamotForbNative
Morus alba L.white mulberryTreeIntroduced
Muhlenbergia schreberi J.F. Gmel.nimblewillGraminoidNative
Oenothera biennis L.common evening primroseForbNative
Oligoneuron rigidum (L.) Smallstiff goldenrodForbNative
Pancium capillare L.witchgrassGraminoidNative
Panicum virgatum L.switchgrassGraminoidNative
Pascopyrum smithii (Rydb.) Á. Lövewestern wheatgrassGraminoidNative
Pediomelum argophyllum (Pursh) J. Grimessilverleaf Indian breadrootLegumeNative
Phalaris arundinacea L.reed canarygrassGraminoidNative
Physalis virginiana Mill.Virginia groundcherryForbNative
Phytolacca americana L.American pokeweedForbNative
Plantago major L.common plantainForbIntroduced
Poa pratensis L.Kentucky bluegrassGraminoidNative
Polygonum arenastrum Jord. ex Boreauoval-leaf knotweedForbIntroduced
Populus deltoides W. Bartram ex Marshall ssp. moniliferaplains cottonwoodTreeNative
Prunus americana MarshallAmerican plumShrubNative
Ratibida columnifera (Nutt.) Woot. & Standl.upright prairie coneflowerForbNative
Rhus glabra L.smooth sumacShrubNative
Rosa arkansana Porterprairie roseShrubNative
Rudbeckia hirta L.blackeyed SusanForbNative
Sagittaria calycina Engelm.hooded arrowheadForb Native
Salsola tragus L.prickly Russian thistleForbIntroduced
Salvia azurea Michx. ex Lam.azure blue sageForbNative
Sambucus nigra L. ssp. canadensis (L.) R. BolliAmerican black elderberryShrubNative
Schizachyrium scoparium (Michx.) Nashlittle bluestemGraminoidNative
Setaria viridis (L.) P. Beauv.green bristlegrassGraminoidIntroduced
Silphium integrifolium Michx.wholeleaf rosinweedForbNative
Silphium laciniatum L.compassplantForbNative
Sisymbrium loeselii L.small tumbleweed mustardForbIntroduced
Solanum rostratum Dunalbuffalobur nightshadeForbNative
Solidago altissima L.Canada goldenrodForbNative
Solidago missouriensis Nutt.Missouri goldenrodForbNative
Sonchus arvensis L.field sowthistleForbIntroduced
Sonchus asper (L.) Hillspiny sowthistleForbIntroduced
Sorghastrum nutans (L.) NashIndiangrassGraminoidNative
Sporobolus heterolepis (A. Gray) A. Grayprairie dropseedGraminoidNative
Symphoricarpos orbiculatus MoenchcoralberryShrubNative
Taraxacum officinale F.H. Wigg.common dandelionForbIntroduced
Tragopogon dubius Scop.yellow salsifyForbIntroduced
Tribulus terrestris L.puncturevineForbIntroduced
Trifolium pratense L.red cloverLegumeIntroduced
Typha angustifolia L.narrowleaf cattailForbNative
Ulmus pumila L.Siberian elmTreeIntroduced
Ulmus rubra Muhl.slippery elmTreeNative
Verbena hastata L.  swamp verbenaForbNative
Vernonia baldwinii Torr.Baldwin’s ironweedForbNative
Vitis riparia Michx.riverbank grapeVineNative

One common grass at this and most sites is “king of native grasses,” big bluestem. It was the main constituent of prairie hay. The Chippewa used big bluestem as medicine for indigestion and stomach pains. The Omaha made a decoction of the the leaf blades to treat general debility from an unknown cause. For fevers, a cut was made on the top of the head and bathed with this decoction. If a fever developed from a wound, lower leaves were applied directly to the wound. The Omaha-Ponca name for big bluestem was hade-zhide, meaning “red hay.” Early settlers found that corn grew best where “big blue” grew. Most of the corn land of the Corn Belt once supported stands of big bluestem.

Pictures of plants found at Site 1: 

The team identified 112 species of the 348 on their plant list, and also found a few not on the checklist, for example Salsola tragus. The trekkers departed for lunch at the Denton Daily Double, and after deliberation returned to the Spring Creek visitor’s center, which had opened by then, where a plant list and even more books were added to the traveling library. 

A note on why the team did not find all 348 species at Spring Creek. The late summer/fall species had asserted themselves, “hiding” most spring and early summer species, and the site was considerably larger than the area sampled.  The group felt confident the plants of the day had introduced themselves, reinforcing the desirability of visiting these prairies often throughout the season.  If not just to clear the mind.

Inventory Site 2 – Nine-Mile Prairie, University of Nebraska, Lancaster County, NE 

This site is owned by University of Nebraska Foundation (purchased in 1983) and leased to the University of Nebraska. Nine-Mile Prairie is a 230-acre relict tallgrass prairie. The prairie was so named because it is five miles west and four miles north of the University campus in downtown Lincoln. Previous botanists have identified 392 vascular plant species. Management consists of springtime burning on a 3-year fire-return interval, along with periodic haying and weed/brush control using herbicides. The prairie has not been grazed since 1968. it serves as a nationally important outdoor laboratory for the study of biological processes in grasslands. 

Nine-Mile is the longest-studied natural area in Nebraska, serving as the site of pioneering research in plant ecology by Professor John E. Weaver, a reputed father of grassland ecology.  The team observed side by side comparison of impacts of burning on reducing the smooth brome component of the prairie.  It is somewhat ironic that smooth brome came to the California Experiment Station in 1884, its promise as a pasture species resulting in seed packets sent to stations in most other states, where it certainly has found a home in the Great Plains.  The team was to have their fill of corn, soybeans, smooth brome, and wind turbines the rest of the week.

Scientific NameCommon NameTypeOrigin
Achillea millefolium L.common yarrowForbNative
Agrostis gigantea RothredtopGraminoidIntroduced
Ambrosia psilostachya DC.Cuman ragweedForbNative
Amorpha canescens PurshleadplantLegumeNative
Andropogon gerardii Vitmanbig bluestemGraminoidNative
Artemisia campestris L.field sagewortForbNative
Artemisia ludoviciana Nutt.white sagebrushForbNative
Ascelpias syriaca L.common milkweedForbNative
Bouteloua curtipendula (Michx.) Torr.sideoats gramaGraminoidNative
Brickellia eupatorioides (L.) Shinnersfalse bonesetForbNative
Bromus inermis Leyss.smooth bromeGraminoidIntroduced
Callirhoe involucrata (Torr. & A. Gray) A. Graypurple poppymallowForbNative
Carex inops L.H. Bailey ssp. heliophilasun sedgeGraminoidNative
Celtis occidentaliseastern hackberryTreeNative
Chloris verticillata Nutt.tumble windmill grassGraminoidNative
Cirsium altissimum (L.) Hilltall thistleForbNative
Convolvulus arvensis L.field bindweedForbIntroduced
Conyza canadensis (L.) CronquistCanadian horseweedForbNative
Cornus drummondii C.A. Mey.roughleaf dogwoodShrubNative
Dalea candida Michx. ex Willd.white prairiecloverLegumeNative
Dalea purpurea Vent.purple prairiecloverLegumeNative
Desmodium illinoense A. GrayIllinois ticktrefoilLegumeNative
Dianthus armeria L.Deptford pinkForbIntroduced
Eragrostis spectabilis (Pursh) Steud.purple lovegrassGraminoidNative
Erigeron strigosus Muhl. ex Willd.prairie fleabaneForbNative
Euphorbia dentata Michx.toothed spurgeForbNative
Euphorbia marginata Purshsnow on the mountainForbNative
Gaura parviflora Douglas ex Lehm.velvetweedForbNative
Helianthus grosseserratus M. Martenssawtooth sunflowerForbNative
Helianthus maximiliani Schrad.Maximillian sunflowerForbNative
Helianthus pauciflorus Nutt. ssp. pauciflorus stff sunflowerForbNative
Heliopsis helianthoides (L.) Sweetsmooth oxeyeForbNative
Koeleria macrantha (Ledeb.) Schult.prairie junegrassGraminoidNative
Lactuca serriola L.prickly lettuceForbIntroduced
Lespedeza capitata Michx.roundhead lespedezaLegumeNative
Liatris aspera Michx.tall blazing starForbNative
Medicago sativa L. alfalfaLegumeIntroduced
Melilotus officinalis (L.) Lam.yellow sweetcloverLegumeIntroduced
Oligoneuron rigidum (L.) Smallstiff goldenrodForbNative
Panicum virgatum L.switchgrassGraminoidNative
Pediomelum argophyllum (Pursh) J. Grimessilverleaf Indian breadrootLegumeNative
Physalis longifolia Nutt.longleaf groundcherryForbNative
Poa pratensis L.Kentucky bluegrassGraminoidNative
Psoralidium lanceolatum (Pursh) Rydb.lemon scurfpeaLegumeNative
Psoralidium tenuiflorum (Pursh) Rydb.slimflower scurfpeaLegumeNative
Rhus glabra L.smooth sumacShrubNative
Rosa arkansana Porterprairie roseShrubNative
Salvia azurea Michx. ex Lam.azure blue sageForbNative
Schizachyrium scoparium (Michx.) Nashlittle bluestemGraminoidNative
Solidago altissima L.Canada goldenrodForbNative
Sorghastrum nutans (L.) NashIndiangrassGraminoidNative
Sporobolus compositus (Poir.) Merr.composite dropseedGraminoidNative
Sporobolus heterolepis (A. Gray) A. Grayprairie dropseedGraminoidNative
Symphoricarpos occidentalis Hook.western snowberryShrubNative
Trifolium pratense L.red cloverLegumeIntroduced
Vernonia baldwinii Torr.Baldwin’s ironweedForbNative

Pictures of plants found at Site 2: 


Having identified and recorded 179 plants on our first day out, we headed north through the corn and soybeans of eastern Nebraska for Sioux City, Iowa, stopping in Wahoo for ice cream.  As Oaks drove on he lobbied with multiple motels at our destination for the best rate, constantly being encouraged to do better by his fellow travelers. The results were satisfactory. 

Day 2 – Monday, August 19, 1913

Departing at 8:00 am, Carlson guided us northeast from Sioux City, Iowa into Plymouth County for the Nature Conservancy’s Broken Kettle Grasslands Preserve. Part of the in-car conversation that morning concerned the red spots and itching on Sharp’s soon-to-be not so private areas. The diagnosis was chiggers.  Sympathy abounded with comments like “You got ‘em, you keep ‘em”. That attitude was short-lived. 

Inventory Site 3 – Broken Kettle Grasslands Preserve, Plymouth County, IA

Broken Kettle remains a sweeping reminder of nature’s beauty. Not only is it the Nature Conservancy’s largest preserve in Iowa but also it contains the largest remaining prairie in Iowa.  In 1999, the Conservancy found the prairie rattlesnake, an extremely rare species, at this site, making Broken Kettle even more important to the region. The Nature Conservancy welcomed a herd of 28 bison at Broken Kettle Grasslands preserve in the fall of 2008.  Broken Kettle Grasslands forms the core of the protection efforts in the northern portion of the Loess Hills and is a stellar example of this rare landform.  Iowa’s Loess Hills were literally blown into existence as the glaciers from the last Ice Age retreated. The extent and depth of the loess deposits along Iowa’s western border are matched only in China. 

Upon arrival at the preserve Carlson conversed with staff at the office and was told they had just released a crop, hatch, herd or whatever, of western rattlesnakes and we might want to avoid the hills immediately bordering the Missouri River flood plain. That seemed reasonable, so Jack led east into the hills, which may well have been the most fruitful and attractive prairie of the entire trek. Smooth brome greeting us at the entrance was a little deceptive, walking up the slope at a good pace we quickly went to work. 

Scientific NameCommon NameTypeOrigin
Acer negundo L.boxelderTreeNative
Agalinis aspera (Douglas ex Benth.) Brittonfalse tall foxgloveForbNative
Ambrosia psilostachya DC.Cuman ragweedForbNative
Amorpha canescens PurshleadplantLegumeNative
Andropogon gerardii Vitmanbig bluestemGraminoidNative
Anemone cylindrica A. Graycandle anemoneForbNative
Artemisia campestris L.field sagewortForbNative
Ascelpias syriaca L.common milkweedForbNative
Bouteloua curtipendula (Michx.) Torr.sideoats gramaGraminoidNative
Bromus inermis Leyss.smooth bromeGraminoidIntroduced
Cannabis sativa L.marijuanaForbIntroduced
Celtis occidentaliseastern hackberryTreeNative
Chamaecrista fasciculata (Michx.) Greenepartridge peaLegumeNative
Chamaesyce glyptosperma (Engelm.) Smallribseed sandmatForbNative
Cirsium flodmanii (Rydb.) ArthurFlodman’s thistleForbNative
Comandra umbellata (L.) Nutt.bastard toadflaxForbNative
Conyza canadensis (L.) CronquistCanadian horseweedForbNative
Cornus drummondii C.A. Mey.roughleaf dogwoodShrubNative
Dalea candida Michx. ex Willd.white prairiecloverLegumeNative
Dalea enneandra Nutt.nineanther prairie cloverLegumeNative
Dalea purpurea Vent.purple prairiecloverLegumeNative
Echinacea angustifolia DC.blacksamson echinaceaForbNative
Elymus canadensis L.Canada wildryeGraminoidNative
Equisetum laevigatum A. Braunsmooth horsestailForbNative
Erigeron strigosus Muhl. ex Willd.prairie fleabaneForbNative
Euphorbia esula L.leafy spurgeForbIntroduced
Euphorbia marginata Purshsnow on the mountainForbNative
Fraxinus pennsylvanica Marshallgreen ashTreeNative
Helianthus annuus L.common sunflowerForbNative
Helianthus pauciflorus Nutt. ssp. pauciflorus stff sunflowerForbNative
Heliopsis helianthoides (L.) Sweetsmooth oxeyeForbNative
Juglans nigra L.black walnutTreeNative
Juniperus virginiana L.eastern redcedarTreeNative
Koeleria macrantha (Ledeb.) Schult.prairie junegrassGraminoidNative
Lactuca tatarica (L.) C.A. Mey.blue lettuceForbNative
Liatris aspera Michx.tall blazing starForbNative
Liatris punctata Hook.dotted blazing starForbNative
Linum sulcatum Riddellgrooved flaxForbNative
Lygodesmia juncea (Pursh) D. Don ex Hook.rush skeletonplantForbNative
Machaeranthera pinnatifida (Hook.) Shinnerslacy tansyasterForbNative
Melilotus officinalis (L.) Lam.yellow sweetcloverLegumeIntroduced
Melilotus officinalis (L.) Lam. (alba)white sweetcloverLegumeIntroduced
Muhlenbergia cuspidata (Torr. ex Hook.) Rydb.plains muhlyGraminoidNative
Oligoneuron rigidum (L.) Smallstiff goldenrodForbNative
Oxytropis lambertii Purshpurple locoweedLegumeNative
Pancium capillare L.witchgrassGraminoidNative
Panicum virgatum L.switchgrassGraminoidNative
Pediomelum argophyllum (Pursh) J. Grimessilverleaf Indian breadrootLegumeNative
Penstemon grandiflorus Nutt.large beardtongueForbNative
Poa palustris L.fowl bluegrassGraminoidNative
Poa pratensis L.Kentucky bluegrassGraminoidNative
Populus deltoides W. Bartram ex Marshall ssp. moniliferaplains cottonwoodTreeNative
Prunus americana MarshallAmerican plumShrubNative
Rhus glabra L.smooth sumacShrubNative
Rosa arkansana Porterprairie roseShrubNative
Schizachyrium scoparium (Michx.) Nashlittle bluestemGraminoidNative
Securigera varia (L.) LassencrownvetchLegumeIntroduced
Setaria viridis (L.) P. Beauv.green bristlegrassGraminoidIntroduced
Solidago altissima L.Canada goldenrodForbNative
Solidago speciosa Nutt.showy goldenrodForbNative
Sorghastrum nutans (L.) NashIndiangrassGraminoidNative
Spartina pectinata Bosc ex Linkprairie cordgrassGraminoidNative
Symphoricarpos occidentalis Hook.western snowberryShrubNative
Verbena hastata L.  swamp verbenaForbNative
Vitis riparia Michx.riverbank grapeVineNative

Lead plant (Amorphia canescens) was found at most sites, including Broken Kettle Grasslands. The Oglala and other American Indians used the dried leaves for smoking and making tea. The tea was also used as a treatment for pinworms. Leaves were steeped and and the liquid was used to wash and treat eczema. The Omaha used a paste made from lead plant for rheumatism and neuralgia. Trek members chewed the leaves in hopes it would ease the chigger bit itching, but it didn’t. 

Pictures of plants found at Site 3: 


After an enjoyable morning and lunch at Bev’s by the (Missouri) River in Sioux City, our next site was right in town.

Inventory Site 4 – Center for Prairie Studies, Briar Cliff University, Sioux City, IA

This 150-acre Nature Conservancy preserve protects the nation’s largest urban tallgrass prairie. Located directly west of campus, it offers Briar Cliff students a noteworthy outdoor classroom. With that in mind, Briar Cliff established the Center for Prairie Studies in 2009, thanks to a major grant from the Gilchrist Foundation. The Center has received additional support from individuals, the Woodbury County Conservation Foundation, the Loess Hills Audubon Society, and the Loess Hills Alliance. The Center provides leadership regarding Briar Cliff’s goal to educate, acquire and respect a sense of place from the prairies of Iowa’s Loess Hills to the global arena.

This site is in the process or reverting to prairie species through management, and Carlson recorded just 50 species.  The site gave the team the first glimpse of fire on the mountain, a miniature of its more famous cousin the Christmas poinsettia. Here it was, hidden down in the robust surrounding vegetation, but not likely to escape the photo lens of Oaks.  We quickly went to work.

Scientific NameCommon NameTypeOrigin
Ambrosia psilostachya DC.Cuman ragweedForbNative
Amorpha canescens PurshleadplantLegumeNative
Andropogon gerardii Vitmanbig bluestemGraminoidNative
Artemisia campestris L.field sagewortForbNative
Artemisia dracunculus L.tarragonForbNative
Artemisia ludoviciana Nutt.white sagebrushForbNative
Asclepias verticillata L.whorled milkweedForb Native
Bouteloua curtipendula (Michx.) Torr.sideoats gramaGraminoidNative
Bromus inermis Leyss.smooth bromeGraminoidIntroduced
Calamovilfa longifolia (Hook.) Scribn.prairie sandreedGraminoidNative
Cannabis sativa L.marijuanaForbIntroduced
Celtis occidentaliseastern hackberryTreeNative
Comandra umbellata (L.) Nutt.bastard toadflaxForbNative
Cornus drummondii C.A. Mey.roughleaf dogwoodShrubNative
Dalea candida Michx. ex Willd.white prairiecloverLegumeNative
Echinacea angustifolia DC.blacksamson echinaceaForbNative
Elymus canadensis L.Canada wildryeGraminoidNative
Eragrostis spectabilis (Pursh) Steud.purple lovegrassGraminoidNative
Euphorbia cyathophora Murrayfire on the mountainForbNative
Euphorbia esula L.leafy spurgeForbIntroduced
Euphorbia marginata Purshsnow on the mountainForbNative
Fraxinus pennsylvanica Marshallgreen ashTreeNative
Helianthus pauciflorus Nutt. ssp. pauciflorus stiff sunflowerForbNative
Heliopsis helianthoides (L.) Sweetsmooth oxeyeForbNative
Juglans nigra L.black walnutTreeNative
Juniperus virginiana L.eastern redcedarTreeNative
Liatris aspera Michx.tall blazing starForbNative
Lonicera tatarica L.Tatarian honeysuckleShrubIntroduced
Lygodesmia juncea (Pursh) D. Don ex Hook.rush skeletonplantForbNative
Machaeranthera pinnatifida (Hook.) Shinnerslacy tansyasterForbNative
Medicago lupulina L.black medickLegumeIntroduced
Melilotus officinalis (L.) Lam.yellow sweetcloverLegumeIntroduced
Melilotus officinalis (L.) Lam. (alba)white sweetcloverLegumeIntroduced
Muhlenbergia cuspidata (Torr. ex Hook.) Rydb.plains muhlyGraminoidNative
Nassella viridula (Trin.) Barkworthgreen needlegrassGraminoidNative
Oligoneuron rigidum (L.) Smallstiff goldenrodForbNative
Onosmodium bejariense DC. ex A. DC.soft-hair marbleseedForbNative
Poa pratensis L.Kentucky bluegrassGraminoidNative
Polygonum arenastrum Jord. ex Boreauoval-leaf knotweedForbIntroduced
Ratibida columnifera (Nutt.) Woot. & Standl.upright prairie coneflowerForbNative
Rhus glabra L.smooth sumacShrubNative
Schizachyrium scoparium (Michx.) Nashlittle bluestemGraminoidNative
Solidago altissima L.Canada goldenrodForbNative
Solidago speciosa Nutt.showy goldenrodForbNative
Spartina pectinata Bosc ex Linkprairie cordgrassGraminoidNative
Sporobolus compositus (Poir.) Merr.composite dropseedGraminoidNative
Sporobolus heterolepis (A. Gray) A. Grayprairie dropseedGraminoidNative
Symphoricarpos occidentalis Hook.western snowberryShrubNative
Verbena hastata L.  swamp verbenaForbNative

The lovely fire on the mountain (Euphorbia cyathophora) (Figure 4-3) is a member of the spurge family. Most members of the family are poisonous. The non-chigger inflicted trek trek members suggested chewing this plant as a cure.

The team experienced a special treat leaving the site, a group photo taken by the practicing university cheerleader squad.

On the road continuing north on I-29, Oaks exited and proceeded west to the east side of the Missouri a few miles to our next, for a non-inventory site.

Adams Homestead and Natural Preserve, Clay County, SD

This stop was more of an educational site, located just north of Sioux City, but in South Dakota. The property contains 1,500 acres along the Missouri River, and was donated to the people of South Dakota in 1984 by Mary and Maud Adams, granddaughters of original homesteader Stephen Searl Adams. They envisioned the area as a place where people, particularly youth, could enjoy the land and learn more about the natural world surrounding them. The site features miles of trails that wind through prairie, forest and along the riverbank. 

With over 100 species of birds, a variety of native plants and a number of different animals, visitors are immersed in nature. Several pioneer period buildings had been mover to the site for preservation. One, a church, had been moved from near the town of Platte, more than 100 miles away and birthplace of Jack’s mother. 

A relaxing stop, and from there the team drove through Sioux Falls and Brookings to Watertown, South Dakota for relatively uneventful evening, except for the discovery of additional infestations of chiggers affecting other members of the team.  The evening meal was taken at Dempsey’s brew pub.

Day 3 – Tuesday, August 20, 1913

Inventory Site 5 – Waubay National Wildlife Refuge, Day County, SD

Waubay National Wildlife Refuge, located in northeastern South Dakota a short drive north from Watertown, was established in 1935 as a refuge and breeding ground for migratory birds and other wildlife. “Waubay,” means “a place where numbers of birds make their nests” in the Dakota language. The Refuge encompasses 4,650 acres of wetlands, native tallgrass prairie, and bur oak forest that provide a wide variety of nesting habitat for more than 100 species of waterfowl, song birds, and upland game birds. Water levels in prairie lakes, like Waubay Lake, are subject to extreme fluctuations. In the 1930’s, Waubay Lake was dry. Heavy precipitation between 1993 and 1997 caused the lake level to rise 15 feet, flooding 100 year old trees.

The office, refuge setting, landscape, and even the vegetation of Waubay provided a diverse experience.  In spite of an overabundance of smooth brome 76 species were identified, including those spotted from the top of a 100-foot tower.  Two inventories were taken:  Waubay prairie and Waubay tower.

The inventory of the prairie at Waubay:

Scientific NameCommon NameTypeOrigin
Allium cernuum Rothnodding onionForbNative
Allium stellatum Fraser ex Ker Gawl.autumn onionForbNative
Amaranthus retroflexus L.redroot amaranthForbNative
Ambrosia psilostachya DC.Cuman ragweedForbNative
Amorpha canescens PurshleadplantLegumeNative
Andropogon gerardii Vitmanbig bluestemGraminoidNative
Anemone cylindrica A. Graycandle anemoneForbNative
Apocynum androsaemifolium L.spreading dogbaneForbNative
Artemisia campestris L.field sagewortForbNative
Artemisia dracunculus L.tarragonForbNative
Artemisia ludoviciana Nutt.white sagebrushForbNative
Ascelpias syriaca L.common milkweedForbNative
Asparagus officinalis L.garden asparagusForb Introduced
Astragalus agrestis Douglas ex G. Donpurple milkvetchLegumeNative
Bouteloua curtipendula (Michx.) Torr.sideoats gramaGraminoidNative
Brickellia eupatorioides (L.) Shinnersfalse bonesetForbNative
Bromus inermis Leyss.smooth bromeGraminoidIntroduced
Cannabis sativa L.marijuanaForbIntroduced
Carex L.sedgeGraminoidNative
Cirsium arvense (L.) Scop.Canada thistleForbIntroduced
Cirsium flodmanii (Rydb.) ArthurFlodman’s thistleForbNative
Cirsium vulgare (Savi) Ten.bull thistleForbIntroduced
Comandra umbellata (L.) Nutt.bastard toadflaxForbNative
Conyza canadensis (L.) CronquistCanadian horseweedForbNative
Crataegus chrysocarpa Ashefireberry hawthornShrubNative
Crataegus succulenta Schrad. ex Linkfleshy hawthornShrubNative
Dalea purpurea Vent.purple prairiecloverLegumeNative
Echinacea angustifolia DC.blacksamson echinaceaForbNative
Elaeagnus angustifolia L.Russian oliveTreeIntroduced
Euphorbia esula L.leafy spurgeForbIntroduced
Galium aparine L.stickywillyForbNative
Glycyrrhiza lepidota PurshAmerican licoriceLegumeNative
Helianthus maximiliani Schrad.Maximillian sunflowerForbNative
Helianthus pauciflorus Nutt. ssp. pauciflorus stiff sunflowerForbNative
Hesperostipa comata (Trin. & Rupr.) Barkworthneedle and threadGraminoidNative
Hordeum jubatum L.foxtail barleyGraminoidNative
Lactuca serriola L.prickly lettuceForbIntroduced
Lactuca tatarica (L.) C.A. Mey.blue lettuceForbNative
Liatris aspera Michx.tall blazing starForbNative
Lonicera dioica L.limber honeysuckleVineNative
Lonicera tatarica L.Tatarian honeysuckleShrubIntroduced
Lygodesmia juncea (Pursh) D. Don ex Hook.rush skeletonplantForbNative
Medicago sativa L. alfalfaLegumeIntroduced
Melilotus officinalis (L.) Lam.yellow sweetcloverLegumeIntroduced
Melilotus officinalis (L.) Lam. (alba)white sweetcloverLegumeIntroduced
Muhlenbergia cuspidata (Torr. ex Hook.) Rydb.plains muhlyGraminoidNative
Oligoneuron rigidum (L.) Smallstiff goldenrodForbNative
Panicum virgatum L.switchgrassGraminoidNative
Pascopyrum smithii (Rydb.) Á. Lövewestern wheatgrassGraminoidNative
Pediomelum argophyllum (Pursh) J. Grimessilverleaf Indian breadrootLegumeNative
Plantago major L.common plantainForbIntroduced
Poa pratensis L.Kentucky bluegrassGraminoidNative
Polygonum arenastrum Jord. ex Boreauoval-leaf knotweedForbIntroduced
Prunus americana MarshallAmerican plumShrubNative
Ratibida columnifera (Nutt.) Woot. & Standl.upright prairie coneflowerForbNative
Rhus glabra L.smooth sumacShrubNative
Rosa arkansana Porterprairie roseShrubNative
Schizachyrium scoparium (Michx.) Nashlittle bluestemGraminoidNative
Shepherdia argentea (Pursh) Nutt.silver buffaloberryShrubNative
Solidago altissima L.Canada goldenrodForbNative
Sonchus arvensis L.field sowthistleForbIntroduced
Sorghastrum nutans (L.) NashIndiangrassGraminoidNative
Spartina pectinata Bosc ex Linkprairie cordgrassGraminoidNative
Symphoricarpos occidentalis Hook.western snowberryShrubNative
Symphyotrichum sericeum (Vent.) G.L. Nesomwestern silver asterForbNative
Taraxacum officinale F.H. Wigg.common dandelionForbIntroduced
Tragopogon dubius Scop.yellow salsifyForbIntroduced
Typha latifolia L.broadleaf cattailForbNative
Vitis riparia Michx.riverbank grapeVineNative

The brief inventory looking down from Waubay tower:

Acer negundo L.boxelderTreeNative
Elaeagnus angustifolia L.Russian oliveTreeIntroduced
Fraxinus pennsylvanica Marshallgreen ashTreeNative
Juniperus virginiana L.eastern redcedarTreeNative
Picea ×albertiana S. Br. [glauca × engelmannii]western white spruceTreeNative
Quercus macrocarpa Michx.bur oakTreeNative
Shepherdia argentea (Pursh) Nutt.silver buffaloberryShrubNative

Purple prairieclover (Dalea purpurea), one of the most beautiful prairie flowers, was medicinally used by American Indians by applying tea made from the leaves to wounds. Ground roots mixed with hot water made a nourishing drink. The Pawnee used the plants and a prophylactic. Early settlers used it the bark of white oak and prairieclover to make a drink used for stopping diarrhea. 




The team departed north from Waubay around lunchtime and enjoyed hot turkey sandwiches with homemade bread at the local café in Eden. Then across the state line through back roads to the next stop at Tewauken National Wildlife Refuge.  During the Civil War, General Sibley’s expedition reached the Dakota Territory and the shore of Lake Tewauken on July 2, 1863.  The expedition included 800 horses, 1,350 wagons, and several hundred head of cattle, often stretching five miles when on the march.  They were in pursuit of a band of Sioux warriers, which had raided settlers in southern Minnesota.  By July 17 Sibley had learned the warriers had left the Devils Lake area and were moving toward the Missouri River. Over the next several days he engaged in three small battles with the band, which escaped across the Missouri. At that point Sibley went back home to Minnesota.  Prior to the Civil War, he was Governor of Minnesota, after the war, president of the Chamber of Commerce in St. Paul.  A little less focused on war than Custer, whose experience in the Dakotas somewhat later led him to Montana to meet his maker.

Inventory Site 6A – Tewaukon National Wildlife Refuge, Sargent County, ND

From the refuge office the team learned that the immediate site might best he inventoried by taking the auto tour around the lake.  The trekkers spotted and recorded 59 species along the way, not counting Zea mays in adjacent fields.

Scientific NameCommon NameTypeOrigin
Acer negundo L.boxelderTreeNative
Amaranthus retroflexus L.redroot amaranthForbNative
Ambrosia psilostachya DC.Cuman ragweedForbNative
Ambrosia trifida L.great ragweedForbNative
Amorpha canescens PurshleadplantLegumeNative
Andropogon gerardii Vitmanbig bluestemGraminoidNative
Apocynum androsaemifolium L.spreading dogbaneForbNative
Artemisia ludoviciana Nutt.white sagebrushForbNative
Ascelpias syriaca L.common milkweedForbNative
Atriplex patula L.spear saltbushForbIntroduced
Bouteloua curtipendula (Michx.) Torr.sideoats gramaGraminoidNative
Bromus inermis Leyss.smooth bromeGraminoidIntroduced
Calamovilfa longifolia (Hook.) Scribn.prairie sandreedGraminoidNative
Caragana arborescens Lam.Siberian peashrubShrubIntroduced
Celtis occidentaliseastern hackberryTreeNative
Chamaesyce glyptosperma (Engelm.) Smallribseed sandmatForbNative
Chenopodium simplex (Torr.) Raf.mapleleaf goosefootForbNative
Cirsium arvense (L.) Scop.Canada thistleForbIntroduced
Convolvulus arvensis L.field bindweedForbIntroduced
Dalea purpurea Vent.purple prairiecloverLegumeNative
Echinacea angustifolia DC.blacksamson echinaceaForbNative
Elaeagnus angustifolia L.Russian oliveTreeIntroduced
Elymus canadensis L.Canada wildryeGraminoidNative
Euphorbia marginata Purshsnow on the mountainForbNative
Fraxinus pennsylvanica Marshallgreen ashTreeNative
Glycyrrhiza lepidota PurshAmerican licoriceLegumeNative
Grindelia squarrosa (Pursh) Dunalcurlycup gumweedForbNative
Helianthus annuus L.common sunflowerForbNative
Helianthus pauciflorus Nutt. ssp. pauciflorus stff sunflowerForbNative
Hordeum jubatum L.foxtail barleyGraminoidNative
Juniperus virginiana L.eastern redcedarTreeNative
Liatris punctata Hook.dotted blazing starForbNative
Lonicera tatarica L.Tatarian honeysuckleShrubIntroduced
Medicago sativa L. alfalfaLegumeIntroduced
Medicago sativa L. ssp. falcata (L.) Arcang.yellow alfalfaLegumeIntroduced
Melilotus officinalis (L.) Lam.yellow sweetcloverLegumeIntroduced
Melilotus officinalis (L.) Lam. (alba)white sweetcloverLegumeIntroduced
Panicum virgatum L.switchgrassGraminoidNative
Pediomelum argophyllum (Pursh) J. Grimessilverleaf Indian breadrootLegumeNative
Phalaris arundinacea L.reed canarygrassGraminoidNative
Phragmites australis (Cav.) Trin. ex Steud.common reedGraminoidNative
Polygonum pensylvanicum L.Pennsylvania smartweedForbNative
Prunus americana MarshallAmerican plumShrubNative
Ratibida columnifera (Nutt.) Woot. & Standl.upright prairie coneflowerForbNative
Rosa arkansana Porterprairie roseShrubNative
Rumex crispus L.curly dockForbIntroduced
Salix amygdaloides Anderssonpeachleaf willowShrubNative
Salix interior Rowleesandbar willowShrubNative
Setaria viridis (L.) P. Beauv.green bristlegrassGraminoidIntroduced
Solidago altissima L.Canada goldenrodForbNative
Sonchus arvensis L.field sowthistleForbIntroduced
Spartina pectinata Bosc ex Linkprairie cordgrassGraminoidNative
Symphyotrichum lanceolatum (Willd.) G.L. Nesomwhite panicle asterForbNative
Taraxacum officinale F.H. Wigg.common dandelionForbIntroduced
Typha latifolia L.broadleaf cattailForbNative
Ulmus pumila L.Siberian elmTreeIntroduced
Verbena hastata L.  swamp verbenaForbNative
Vitis riparia Michx.riverbank grapeVineNative
Xanthium strumarium L.rough cockleburForbNative

Plains Indians favored the medicinal value of blacksamson echinacea, also called purple coneflower (Echinacea angustifolia) very high. It was used for snakebites, bee stings, headaches, stomach cramps, toothaches, enlarged glands such as mumps, sore throats, and hydrophobia and for distemper in horses. The Sioux used freshly scraped root for snakebite and for the bite of a mad dog, and they chewed pieces of root stalk for toothache. They also discovered that it was somewhat like a burn preventative and enabled the body to endure extreme heat. Medicine men bathed their hands and arms in the juice, and then picked out meat from boiling water. Sometimes it was used prior to sweat baths and ritual feats such as immersing hands in scalding water or holding live coals in the mouth. A smoke treatment from the plant was used for headache in humans and for distemper in horses. Modern medicine still uses extracts from this plant for a variety of homeopathic treatments. 


From there Oaks drove a little north to another Tewaukon site for the conventional inventory. 

Inventory Site 6B – Tewaukon National Wildlife Refuge Waterfowl Production Area, Sargent County, ND

The site bordered a pothole, which afforded an introduction to plants on somewhat more saline conditions near the shore.

Scientific NameCommon NameTypeOrigin
Achillea millefolium L.common yarrowForbNative
Allium stellatum Fraser ex Ker Gawl.autumn onionForbNative
Ambrosia psilostachya DC.Cuman ragweedForbNative
Amorpha canescens PurshleadplantLegumeNative
Amorpha fruticosa L.false indigo bushShrubNative
Andropogon gerardii Vitmanbig bluestemGraminoidNative
Apocynum androsaemifolium L.spreading dogbaneForbNative
Artemisia ludoviciana Nutt.white sagebrushForbNative
Ascelpias incarnata L.swamp milkweedForbNative
Ascelpias syriaca L.common milkweedForbNative
Bassia scoparia (L.) A.J. ScottburningbushForbIntroduced
Bouteloua curtipendula (Michx.) Torr.sideoats gramaGraminoidNative
Bromus inermis Leyss.smooth bromeGraminoidIntroduced
Carex L.sedgeGraminoidNative
Cirsium arvense (L.) Scop.Canada thistleForbIntroduced
Cirsium vulgare (Savi) Ten.bull thistleForbIntroduced
Conyza canadensis (L.) CronquistCanadian horseweedForbNative
Dichanthelium oligosanthes (Schult.) GouldScribner’s rosette grassGraminoidNative
Echinacea angustifolia DC.blacksamson echinaceaForbNative
Echinochloa crus-galli (L.) P. Beauv.barnyardgrassGraminoidIntroduced
Elaeagnus angustifolia L.Russian oliveTreeIntroduced
Erigeron strigosus Muhl. ex Willd.prairie fleabaneForbNative
Glycyrrhiza lepidota PurshAmerican licoriceLegumeNative
Helianthus maximiliani Schrad.Maximillian sunflowerForbNative
Hordeum jubatum L.foxtail barleyGraminoidNative
Lycopus americanus Muhl. ex W.P.C. BartonAmerican water horehoundForbNative
Medicago lupulina L.black medickLegumeIntroduced
Medicago sativa L. alfalfaLegumeIntroduced
Melilotus officinalis (L.) Lam.yellow sweetcloverLegumeIntroduced
Nassella viridula (Trin.) Barkworthgreen needlegrassGraminoidNative
Oligoneuron rigidum (L.) Smallstiff goldenrodForbNative
Onosmodium bejariense DC. ex A. DC.soft-hair marbleseedForbNative
Pancium capillare L.witchgrassGraminoidNative
Pascopyrum smithii (Rydb.) Á. Lövewestern wheatgrassGraminoidNative
Pediomelum argophyllum (Pursh) J. Grimessilverleaf Indian breadrootLegumeNative
Phalaris arundinacea L.reed canarygrassGraminoidNative
Phragmites australis (Cav.) Trin. ex Steud.common reedGraminoidNative
Poa pratensis L.Kentucky bluegrassGraminoidNative
Polygonum pensylvanicum L.Pennsylvania smartweedForbNative
Ratibida columnifera (Nutt.) Woot. & Standl.upright prairie coneflowerForbNative
Rosa arkansana Porterprairie roseShrubNative
Rumex crispus L.curly dockForbIntroduced
Schizachyrium scoparium (Michx.) Nashlittle bluestemGraminoidNative
Scirpus pallidus (Britton) Fernaldcloaked bulrushGraminoidNative
Solidago altissima L.Canada goldenrodForbNative
Sonchus arvensis L.field sowthistleForbIntroduced
Spartina pectinata Bosc ex Linkprairie cordgrassGraminoidNative
Symphoricarpos occidentalis Hook.western snowberryShrubNative
Symphyotrichum ciliatum (Ledeb.) G.L. Nesomrayless alkali asterForbNative
Vernonia fasciculata Michx.prairie ironweedForbNative

In 1949, E.J. Dyksterhuis proposed a quantitative system for assessing whether a rangeland was at an early or late stage of succession by analyzing the behaviors of three classes of plant species; decreasers, increasers, and invaders. The ecologists and the rancher thought late stages were the most desirable. However, livestock grazing drove the plant composition toward earlier stages, causing late stage species to decrease in abundance. These decreasers were replaced by other plants that increased in abundance, thus called increasers. Dyksterhuis also proposed that the condition of rangelands improved as succession progressed toward later stages, which provide better forage and to be more stable and productive plant communities. The condition of a rangeland site could therefore be determined by its status between early and late stage succession. 

Unfortunately, decreasers are few compared to the number of increaser. The Tewaukon site is typical. The best decreaser species are the native tall grasses. This site contains ruminants of most, if not all of them. But their percent of the total forage component is low. As disturbances, such as over grazing, takes place, both native and introduced increases invade. For example, stiff goldenrod (Oligoneuron rigidum) is an invader in over grazed rangeland, even though it is a native and provides lovely color to the fall rangeland. 

Inventorying for the day was done. Oaks took his last picture at 5:18 PM, and then headed north to Lisbon, 40 miles away.  

On the way Carlson informed fellow trekkers the van was descending the Coteau des Prairies, or Prairie Coteau, dropping 900 feet in elevation into the Red River valley. The Coteau is the prominent, wedge-shaped landform whose point just juts into southeastern North Dakota. Although the Coteau is cored by bedrock, outcrops of bedrock are limited.  Instead the feature is draped by glacial deposits whose thickness averages over 400 feet. The view in the picture below is as if one were flying south from the region of Geneseo into South Dakota. The brown state line is visible; the picture exaggerates vertical elevation by 20:1.

Day 4 – Wednesday, August 21, 1913

The Sheyenne National Grassland operated by the U.S. Forest Service (USFS) comprises approximately 70,180 acres mostly in Ransom County, North Dakota, scattered over many sections.  A visit to the USFS office directed the team first to the ‘most pristine’ section, believed never to have been grazed.  The next inventory was conducted at what the record keeper called Pigeon Point in the north central part of the Grassland.  Following lunch in nearby downtown Sheldon at D’s Dear Island café (more hot turkey sandwiches and a peanut butter dutter burger) the team turned east to the third site, labeled ‘Unquestionably Pristine’. 

The Grassland contains the largest population of the greater prairie chicken in North Dakota.  Of 1,200 plant species in North Dakota, 850 can be found on the Grassland, such as the threatened western prairie white-fringed orchid and beach heather.  Soils are mostly sandy, geologically a former delta of an old river draining into the Pleistocene Lake Agassiz, today supporting prairie and not corn and soybeans growing on the more loamy soils surrounding the Grassland

Ransom County is thinly populated. In 1800 they had 527 souls; by 1920 it was 11,618.  The most recent census says 5,444, not counting botanists.  That works out to about 7 people per square mile, and while the plant population is at least moderately diverse the same cannot be said about the human population. The demographic makeup of the county is 97.93% white, made up of 39.5% German and 34.9% Norwegian ancestry. 

Inventory Site 7A – Sheyenne National Grassland, Ransom County, ND, Ungrazed

Oaks drove the team to the first site in the southeast part of the grassland, through a pasture gate, knee high grass, and a few fragrant cow pies until reaching the ungrazed lot. 

Scientific NameCommon NameTypeOrigin
Allium stellatum Fraser ex Ker Gawl.autumn onionForbNative
Amorpha canescens PurshleadplantLegumeNative
Andropogon gerardii Vitmanbig bluestemGraminoidNative
Anemone canadensis L.Canada anemoneForbNative
Apocynum androsaemifolium L.spreading dogbaneForbNative
Artemisia ludoviciana Nutt.white sagebrushForbNative
Ascelpias syriaca L.common milkweedForbNative
Asclepias verticillata L.whorled milkweedForb Native
Bouteloua curtipendula (Michx.) Torr.sideoats gramaGraminoidNative
Bromus inermis Leyss.smooth bromeGraminoidIntroduced
Calamagrostis stricta (Timm) Koelerslimstem reedgrassGraminoidNative
Carex L.sedgeGraminoidNative
Cirsium flodmanii (Rydb.) ArthurFlodman’s thistleForbNative
Convolvulus arvensis L.field bindweedForbIntroduced
Dalea candida Michx. ex Willd.white prairiecloverLegumeNative
Dalea purpurea Vent.purple prairiecloverLegumeNative
Echinacea angustifolia DC.blacksamson echinaceaForbNative
Elymus canadensis L.Canada wildryeGraminoidNative
Equisetum laevigatum A. Braunsmooth horsestailForbNative
Euphorbia esula L.leafy spurgeForbIntroduced
Glycyrrhiza lepidota PurshAmerican licoriceLegumeNative
Grindelia squarrosa (Pursh) Dunalcurlycup gumweedForbNative
Helianthus maximiliani Schrad.Maximillian sunflowerForbNative
Lactuca serriola L.prickly lettuceForbIntroduced
Liatris aspera Michx.tall blazing starForbNative
Liatris punctata Hook.dotted blazing starForbNative
Medicago lupulina L.black medickLegumeIntroduced
Mentha arvensis L.wild mintForbNative
Oligoneuron rigidum (L.) Smallstiff goldenrodForbNative
Panicum virgatum L.switchgrassGraminoidNative
Poa pratensis L.Kentucky bluegrassGraminoidNative
Ratibida columnifera (Nutt.) Woot. & Standl.upright prairie coneflowerForbNative
Rosa arkansana Porterprairie roseShrubNative
Rudbeckia hirta L.blackeyed SusanForbNative
Salix discolor Muhl.pussy willowShrubNative
Salix interior Rowleesandbar willowShrubNative
Schizachyrium scoparium (Michx.) Nashlittle bluestemGraminoidNative
Solidago altissima L.Canada goldenrodForbNative
Sonchus arvensis L.field sowthistleForbIntroduced
Sorghastrum nutans (L.) NashIndiangrassGraminoidNative
Spartina pectinata Bosc ex Linkprairie cordgrassGraminoidNative
Spiraea alba Du Roiwhite meadowsweetShrubNative
Symphoricarpos occidentalis Hook.western snowberryShrubNative
Symphyotrichum ericoides (L.) G.L. Nesomwhite heath asterForbNative
Symphyotrichum lanceolatum (Willd.) G.L. Nesomwhite panicle asterForbNative
Zizia aptera (A. Gray) Fernaldmeadow ziziaForbNative


Inventory Site 7B – Sheyenne National Grassland, Ransom County, ND, Pigeon Point

Site 7A down, the group then trekked north on gravel roads, arriving at a gate to corrals and cattle chutes, a reasonable point for egress into the second recommended site.  The paddock was a landscape of swales and sandy ridges, a variety of seasonably wet and arid habitats, waste high grass in the swales.

Scientific NameCommon NameTypeOrigin
Achillea millefolium L.common yarrowForbNative
Agrostis scabra Willd.rough bentgrassGraminoidNative
Allium stellatum Fraser ex Ker Gawl.autumn onionForbNative
Amaranthus retroflexus L.redroot amaranthForbNative
Ambrosia psilostachya DC.Cuman ragweedForbNative
Amorpha canescens PurshleadplantLegumeNative
Andropogon gerardii Vitmanbig bluestemGraminoidNative
Andropogon hallii Hack.sand bluestemGraminoidNative
Anemone canadensis L.Canada anemoneForbNative
Artemisia campestris L.field sagewortForbNative
Artemisia frigida Willd.prairie sagewortForbNative
Artemisia ludoviciana Nutt.white sagebrushForbNative
Ascelpias syriaca L.common milkweedForbNative
Asclepias verticillata L.whorled milkweedForb Native
Astragalus cicer L.chickpea milkvetchLegumeIntroduced
Bassia scoparia (L.) A.J. ScottburningbushForbIntroduced
Bouteloua gracilis (Willd. ex Kunth) Lag. ex Griffithsblue gramaGraminoidNative
Bromus inermis Leyss.smooth bromeGraminoidIntroduced
Calamovilfa longifolia (Hook.) Scribn.prairie sandreedGraminoidNative
Carex L.sedgeGraminoidNative
Carex praegracilis W. Boottclustered field sedgeGraminoidNative
Chenopodium album L.lambsquartersForbIntroduced
Cirsium arvense (L.) Scop.Canada thistleForbIntroduced
Cirsium flodmanii (Rydb.) ArthurFlodman’s thistleForbNative
Conyza canadensis (L.) CronquistCanadian horseweedForbNative
Dalea candida Michx. ex Willd.white prairiecloverLegumeNative
Dalea purpurea Vent.purple prairiecloverLegumeNative
Daucus carota L.Queen Anne’s laceForbIntroduced
Echinochloa crus-galli (L.) P. Beauv.barnyardgrassGraminoidIntroduced
Elymus canadensis L.Canada wildryeGraminoidNative
Elymus repens (L.) GouldquackgrassGraminoidIntroduced
Eragrostis cilianensis (All.) Vign. ex JanchenstinkgrassGraminoidIntroduced
Eragrostis spectabilis (Pursh) Steud.purple lovegrassGraminoidNative
Erysimum inconspicuum (S. Watson) MacMill.shy wallflowerForbNative
Eupatorium perfoliatum L.common bonesetForbNative
Euphorbia esula L.leafy spurgeForbIntroduced
Helenium autumnale L.common sneezeweedForbNative
Helianthus annuus L.common sunflowerForbNative
Helianthus maximiliani Schrad.Maximillian sunflowerForbNative
Hesperostipa comata (Trin. & Rupr.) Barkworthneedle and threadGraminoidNative
Heterotheca villosa (Pursh) Shinnershairy false goldenasterForbNative
Hordeum jubatum L.foxtail barleyGraminoidNative
Koeleria macrantha (Ledeb.) Schult.prairie junegrassGraminoidNative
Liatris aspera Michx.tall blazing starForbNative
Liatris punctata Hook.dotted blazing starForbNative
Lygodesmia juncea (Pursh) D. Don ex Hook.rush skeletonplantForbNative
Medicago lupulina L.black medickLegumeIntroduced
Melilotus officinalis (L.) Lam. (alba)white sweetcloverLegumeIntroduced
Mirabilis hirsuta (Pursh) MacMill.hairy four o’clockForbNative
Pancium capillare L.witchgrassGraminoidNative
Panicum virgatum L.switchgrassGraminoidNative
Pascopyrum smithii (Rydb.) Á. Lövewestern wheatgrassGraminoidNative
Penstemon grandiflorus Nutt.large beardtongueForbNative
Phalaris arundinacea L.reed canarygrassGraminoidNative
Physalis virginiana Mill.Virginia groundcherryForbNative
Poa pratensis L.Kentucky bluegrassGraminoidNative
Polygonum pensylvanicum L.Pennsylvania smartweedForbNative
Populus deltoides W. Bartram ex Marshall ssp. moniliferaplains cottonwoodTreeNative
Potentilla norvegica L.Norwegian cinquefoilForbNative
Ratibida columnifera (Nutt.) Woot. & Standl.upright prairie coneflowerForbNative
Rosa arkansana Porterprairie roseShrubNative
Rumex crispus L.curly dockForbIntroduced
Salix amygdaloides Anderssonpeachleaf willowShrubNative
Salix interior Rowleesandbar willowShrubNative
Schizachyrium scoparium (Michx.) Nashlittle bluestemGraminoidNative
Schoenoplectus tabernaemontani (C.C. Gmel.) Pallasoftstem bulrushGraminoidNative
Scirpus pallidus (Britton) Fernaldcloaked bulrushGraminoidNative
Setaria viridis (L.) P. Beauv.green bristlegrassGraminoidIntroduced
Solidago altissima L.Canada goldenrodForbNative
Sonchus arvensis L.field sowthistleForbIntroduced
Spartina pectinata Bosc ex Linkprairie cordgrassGraminoidNative
Spiraea alba Du Roiwhite meadowsweetShrubNative
Sporobolus cryptandrus (Torr.) A. Graysand dropseedGraminoidNative
Symphyotrichum lanceolatum (Willd.) G.L. Nesomwhite panicle asterForbNative
Tradescantia occidentalis (Britton) Smythprairie spiderwortForbNative
Trifolium pratense L.red cloverLegumeIntroduced
Trifolium repens L.white cloverLegumeIntroduced
Typha latifolia L.broadleaf cattailForbNative
Verbena hastata L.  swamp verbenaForbNative
Vernonia fasciculata Michx.prairie ironweedForbNative

Inventory Site 7C – Sheyenne National Grassland, Richland County, ND, Unquestionably Pristine

After fruitfully identifying 79 species at the second site, the group took lunch in Sheldon, and then proceeded to the third location at the eastern edge of the Grassland.  Vegetation here was less diverse as the landscape was more uniform, 52 species identified.  This day big bluestem and white sweetclover dominated the view at this site.

Scientific NameCommon NameTypeOrigin
Agrostis scabra Willd.rough bentgrassGraminoidNative
Andropogon gerardii Vitmanbig bluestemGraminoidNative
Anemone canadensis L.Canada anemoneForbNative
Anemone cylindrica A. Graycandle anemoneForbNative
Artemisia campestris L.field sagewortForbNative
Artemisia ludoviciana Nutt.white sagebrushForbNative
Ascelpias incarnata L.swamp milkweedForbNative
Ascelpias syriaca L.common milkweedForbNative
Bromus inermis Leyss.smooth bromeGraminoidIntroduced
Carex L.sedgeGraminoidNative
Chamaesyce glyptosperma (Engelm.) Smallribseed sandmatForbNative
Chenopodium pratericola Rydb.desert goosefootForbNative
Cirsium altissimum (L.) Hilltall thistleForbNative
Cornus sericea L.redosier dogwoodShrubNative
Dalea candida Michx. ex Willd.white prairiecloverLegumeNative
Dalea purpurea Vent.purple prairiecloverLegumeNative
Echinacea angustifolia DC.blacksamson echinaceaForbNative
Equisetum laevigatum A. Braunsmooth horsestailForbNative
Euphorbia esula L.leafy spurgeForbIntroduced
Euthamia graminifolia (L.) Nutt.flat-top goldentopForbNative
Filipendula rubra (Hill) B.L. Rob.queen of the prairieForbNative
Fragaria virginiana DuchesneVirginia strawberryForbNative
Gentiana andrewsii Griseb.closed bottle gentianForbNative
Helianthus annuus L.common sunflowerForbNative
Lycopus americanus Muhl. ex W.P.C. BartonAmerican water horehoundForbNative
Melilotus officinalis (L.) Lam.yellow sweetcloverLegumeIntroduced
Melilotus officinalis (L.) Lam. (alba)white sweetcloverLegumeIntroduced
Pancium capillare L.witchgrassGraminoidNative
Panicum virgatum L.switchgrassGraminoidNative
Phalaris arundinacea L.reed canarygrassGraminoidNative
Poa pratensis L.Kentucky bluegrassGraminoidNative
Populus deltoides W. Bartram ex Marshall ssp. moniliferaplains cottonwoodTreeNative
Potentilla arguta Purshtall cinquefoilForbNative
Prunus virginiana L.chokecherryShrubNative
Pycnanthemum virginianum (L.) T. Dur. & B.D. Jacks. ex B.L. Rob. & FernaldVirginia mountainmintForbNative
Rosa arkansana Porterprairie roseShrubNative
Salix humilis Marshallprairie willowShrubNative
Salix interior Rowleesandbar willowShrubNative
Salix serissima (L.H. Bailey) Fernaldautumn willowShrubNative
Schizachyrium scoparium (Michx.) Nashlittle bluestemGraminoidNative
Scirpus pallidus (Britton) Fernaldcloaked bulrushGraminoidNative
Silphium perfoliatum L.cup plantForbNative
Solidago altissima L.Canada goldenrodForbNative
Sorghastrum nutans (L.) NashIndiangrassGraminoidNative
Spartina pectinata Bosc ex Linkprairie cordgrassGraminoidNative
Spiraea alba Du Roiwhite meadowsweetShrubNative
Symphoricarpos occidentalis Hook.western snowberryShrubNative
Symphyotrichum ericoides (L.) G.L. Nesomwhite heath asterForbNative
Symphyotrichum lanceolatum (Willd.) G.L. Nesomwhite panicle asterForbNative
Trifolium pratense L.red cloverLegumeIntroduced
Typha latifolia L.broadleaf cattailForbNative
Zizia aptera (A. Gray) Fernaldmeadow ziziaForbNative

Sideoats grama (Bouteloua curtipendula), which was present at this and most sites, was the first of the tall grasses to be domesticated and sold as a commercial variety. The first was ‘Vaughn’, named for the location in New Mexico where it was collected.  Commercial seed production from nine varieties of sideoats from 1977-2005 exceeded 10 million pounds. This is similar to the production from switchgrass, another native tallgrass prairie species. Both are decreasers on ruminant prairies.

At day’s end, the team drove north and across the Red River from Fargo to Moorhead, Minnesota, where some serious attention was devoted to chigger cure.  

Day 5 – Thursday, August 22, 1913

While the itching continued there were no reports of new chigger bites. 

After breakfast the team drove east about 17 miles to a Nature Conservancy site. While not apparent the van traveled across the old shoreline of Lake Agassiz, an immense glacial lake located in the middle of the northern part of North America.  Fed by melt water at the end of the last glacial period, its area was larger than all of the modern Great Lakes combined, and at times it held more water than contained by all lakes in the world today.  Several modern river valleys, including the Minnesota River, were originally cut by water entering or leaving the lake. The fertile soils of the valley, now drained by the Red River of the North, are formed from lacustrine deposits of silt from Lake Agassiz.

Inventory Site 8 – Bluestem Prairie Scientific and Natural Area, Clay County, MN

Despite a history of ownership by farmers, this preserve has remained mostly native grassland. Today, it is recognized as one of the largest and highest quality northern tallgrass prairies in the U.S.  When nearby Buffalo River State Park was established, it included only a small portion of native prairie and was used mainly for public recreation along the river. The park boundaries were enlarged significantly in the 1960’s when concern grew over rapidly diminishing grassland. Today, the Conservancy is taking many active steps to ensure the protection of this increasingly rare natural treasure. 

The Conservancy manages the preserve through prescribed burning, biological inventories, and controlling non-native species by hand-cutting, mowing and localized spraying. One of the most troublesome exotic plants is leafy spurge. In recent years the Conservancy restored more than 677 acres of prairie which formerly had been ditched, farmed, and used as a gravel pit.

Scientific NameCommon NameTypeOrigin
Achillea millefolium L.common yarrowForbNative
Agrostis scabra Willd.rough bentgrassGraminoidNative
Allium stellatum Fraser ex Ker Gawl.autumn onionForbNative
Ambrosia psilostachya DC.Cuman ragweedForbNative
Amorpha canescens PurshleadplantLegumeNative
Amorpha fruticosa L.false indigo bushShrubNative
Andropogon gerardii Vitmanbig bluestemGraminoidNative
Anemone cylindrica A. Graycandle anemoneForbNative
Antennaria neglecta Greenefield pussytoesForbNative
Artemisia campestris L.field sagewortForbNative
Artemisia frigida Willd.prairie sagewortForbNative
Artemisia ludoviciana Nutt.white sagebrushForbNative
Ascelpias syriaca L.common milkweedForbNative
Bouteloua curtipendula (Michx.) Torr.sideoats gramaGraminoidNative
Bouteloua gracilis (Willd. ex Kunth) Lag. ex Griffithsblue gramaGraminoidNative
Bromus inermis Leyss.smooth bromeGraminoidIntroduced
Calylophus serrulatus (Nutt.) P.H. Ravenyellow sundropsForbNative
Carex L.sedgeGraminoidNative
Cerastium arvense L.field chickweedForbIntroduced
Chenopodium pratericola Rydb.desert goosefootForbNative
Cirsium flodmanii (Rydb.) ArthurFlodman’s thistleForbNative
Comandra umbellata (L.) Nutt.bastard toadflaxForbNative
Conyza canadensis (L.) CronquistCanadian horseweedForbNative
Dalea candida Michx. ex Willd.white prairiecloverLegumeNative
Dalea purpurea Vent.purple prairiecloverLegumeNative
Daucus carota L.Queen Anne’s laceForbIntroduced
Dichanthelium oligosanthes (Schult.) GouldScribner’s rosette grassGraminoidNative
Echinacea angustifolia DC.blacksamson echinaceaForbNative
Elymus virginicus L.Virginia wildryeGraminoidNative
Equisetum hyemale L.scouringrush horsetailForbNative
Eragrostis spectabilis (Pursh) Steud.purple lovegrassGraminoidNative
Erigeron strigosus Muhl. ex Willd.prairie fleabaneForbNative
Gaillardia aristata PurshblanketflowerForbNative
Galium boreale L.northern bedstrawForbNative
Helianthus petiolaris Nutt.prairie sunflowerForbNative
Hesperostipa comata (Trin. & Rupr.) Barkworthneedle and threadGraminoidNative
Hesperostipa spartea (Trin.) BarkworthporcupinegrassGraminoidNative
Heterotheca villosa (Pursh) Shinnershairy false goldenasterForbNative
Lactuca tatarica (L.) C.A. Mey.blue lettuceForbNative
Liatris aspera Michx.tall blazing starForbNative
Liatris punctata Hook.dotted blazing starForbNative
Linum sulcatum Riddellgrooved flaxForbNative
Lygodesmia juncea (Pursh) D. Don ex Hook.rush skeletonplantForbNative
Medicago lupulina L.black medickLegumeIntroduced
Melilotus officinalis (L.) Lam. (alba)white sweetcloverLegumeIntroduced
Mirabilis hirsuta (Pursh) MacMill.hairy four o’clockForbNative
Muhlenbergia cuspidata (Torr. ex Hook.) Rydb.plains muhlyGraminoidNative
Oenothera biennis L.common evening primroseForbNative
Oligoneuron rigidum (L.) Smallstiff goldenrodForbNative
Onosmodium bejariense DC. ex A. DC.soft-hair marbleseedForbNative
Parthenocissus vitacea (Knerr) Hitchc.woodbineVineNative
Pediomelum argophyllum (Pursh) J. Grimessilverleaf Indian breadrootLegumeNative
Penstemon grandiflorus Nutt.large beardtongueForbNative
Physalis virginiana Mill.Virginia groundcherryForbNative
Poa pratensis L.Kentucky bluegrassGraminoidNative
Prunus pumila L.sandcherryShrubNative
Prunus virginiana L.chokecherryShrubNative
Pycnanthemum virginianum (L.) T. Dur. & B.D. Jacks. ex B.L. Rob. & FernaldVirginia mountainmintForbNative
Ratibida columnifera (Nutt.) Woot. & Standl.upright prairie coneflowerForbNative
Rosa arkansana Porterprairie roseShrubNative
Salix humilis Marshallprairie willowShrubNative
Salix interior Rowleesandbar willowShrubNative
Schizachyrium scoparium (Michx.) Nashlittle bluestemGraminoidNative
Solidago altissima L.Canada goldenrodForbNative
Sorghastrum nutans (L.) NashIndiangrassGraminoidNative
Sporobolus compositus (Poir.) Merr.composite dropseedGraminoidNative
Sporobolus heterolepis (A. Gray) A. Grayprairie dropseedGraminoidNative
Symphoricarpos occidentalis Hook.western snowberryShrubNative
Symphyotrichum lanceolatum (Willd.) G.L. Nesomwhite panicle asterForbNative
Tradescantia occidentalis (Britton) Smythprairie spiderwortForbNative
Trientalis borealis Raf.starflowerForbNative
Vernonia fasciculata Michx.prairie ironweedForbNative
Vitis riparia Michx.riverbank grapeVineNative
Zizia aptera (A. Gray) Fernaldmeadow ziziaForbNative

Leaving this most pleasant site after identifying 74 species, the team continued east to Detroit Lakes, Minnesota and enjoyed lunch on the lakefront with long time colleague Erling Jacobson and his wife Lois.  Also joining was retired SCS range conservationists John Gunderson.  While Erling’s interests are typical of many retirees, an avid fisherman, he maintains enthusiasm for the plant materials program.  It is safe to say that all enjoyed a fresh and uplifting luncheon experience.  Walleye continued to be on menus, here a nice sandwich of fried fish.

Around 2:00 pm Oaks pointed the van south for about 200 miles to Pipestone, MN and the next inventory site at Pipestone in the southwest corner of the state.  On the way some dreamed they were going through Kansas.  Others noted the increase in density of wind turbines.

Securing lodging by way of the Oaks method, the team took their evening meal at the Calumet Inn, the exterior made of various shades of Sioux quartzite from local quarries.

Day 6 – Friday, August 23, 1913

Inventory Site 9 – Pipestone National Monument, Pipestone County, MN

This was a most enjoyable site.  All have seen, many times no doubt, products used in movies that theoretically came from this location. For many generations, Native Americans have quarried the red pipestone found at this site. These grounds are sacred to many people because the pipestone quarried here is carved into pipes used for prayer, peace, healing, ritual dancing, trade, warfare, and other ceremonies. The soft red pipestone is found in a vein between layers of the harder red Sioux quartzite. Methods of quarrying have changed little since the process began. http://www.highhopesgardens.com/Blogphotos/2009/pipestonesign.jpg

Quarrying is a laborious task involving weeks of work with hand tools, including sledgehammers, pry bars, sharp chisels, and metal wedges. Many believe that the pipe’s smoke from this site carries one’s prayer to the Great Spirit. The traditions of quarrying and pipemaking continue here today. 

Additionally, Pipestone National Monument is another of the few remaining areas of native tallgrass prairie. Over 400,000 square miles of tall grass prairie once covered the Midwest. Less than 1% of the original remains today. Pipestone National Monument is located on the same Coteau des Prairies discussed earlier, also called the Highland of the Prairies. The dominant plant communities at the Monument include; virgin native prairie, restored prairie, degraded prairie, and oak savanna. The 301 acre Monument is bisected by several features: the pipestone quarries, a Sioux quartzite cliff line, and Pipestone Creek.

The inventory was preformed from a path winding through the Monument, offering many habitats and picturesque settings.  The team identified 127 species, the most of any site on the trek.

Scientific NameCommon NameTypeOrigin
Acer negundo L.boxelderTreeNative
Achillea millefolium L.common yarrowForbNative
Allium stellatum Fraser ex Ker Gawl.autumn onionForbNative
Ambrosia psilostachya DC.Cuman ragweedForbNative
Ambrosia trifida L.great ragweedForbNative
Amorpha canescens PurshleadplantLegumeNative
Amorpha fruticosa L.false indigo bushShrubNative
Andropogon gerardii Vitmanbig bluestemGraminoidNative
Anemone cylindrica A. Graycandle anemoneForbNative
Apocynum androsaemifolium L.spreading dogbaneForbNative
Arctium minus Bernh.lesser burdockForbIntroduced
Artemisia campestris L.field sagewortForbNative
Artemisia dracunculus L.tarragonForbNative
Artemisia frigida Willd.prairie sagewortForbNative
Ascelpias syriaca L.common milkweedForbNative
Asclepias verticillata L.whorled milkweedForb Native
Asparagus officinalis L.garden asparagusForb Introduced
Bouteloua curtipendula (Michx.) Torr.sideoats gramaGraminoidNative
Bouteloua dactyloides (Nutt.) J.T. ColumbusbuffalograssGraminoidNative
Bromus inermis Leyss.smooth bromeGraminoidIntroduced
Carex filifolia Nutt.threadleaf sedgeGraminoidNative
Carex L.sedgeGraminoidNative
Carum carvi L.carawayForbNative
Celastrus scandens L.American bittersweetVineNative
Celtis occidentaliseastern hackberryTreeNative
Chamaesyce glyptosperma (Engelm.) Smallribseed sandmatForbNative
Chenopodium album L.lambsquartersForbIntroduced
Chloris verticillata Nutt.tumble windmill grassGraminoidNative
Cirsium arvense (L.) Scop.Canada thistleForbIntroduced
Comandra umbellata (L.) Nutt.bastard toadflaxForbNative
Conyza canadensis (L.) CronquistCanadian horseweedForbNative
Cornus sericea L.redosier dogwoodShrubNative
Crataegus chrysocarpa Ashefireberry hawthornShrubNative
Crepis runcinata (James) Torr. & A. Grayfiddleleaf hawksbeardForbNative
Cystopteris tenuis (Michx.) Desv.upland brittle bladderfernForbNative
Echinacea angustifolia DC.blacksamson echinaceaForbNative
Echinochloa crus-galli (L.) P. Beauv.barnyardgrassGraminoidIntroduced
Echinocystis lobata (Michx.) Torr. & A. Graywild cucumberVineNative
Elymus canadensis L.Canada wildryeGraminoidNative
Elymus repens (L.) GouldquackgrassGraminoidIntroduced
Elymus virginicus L.Virginia wildryeGraminoidNative
Erysimum cheiranthoides L.wormseed wallflowerForbNative
Erysimum inconspicuum (S. Watson) MacMill.shy wallflowerForbNative
Euphorbia cyathophora Murrayfire on the mountainForbNative
Euphorbia dentata Michx.toothed spurgeForbNative
Festuca rubra L.red fescueGraminoidIntroduced
Fraxinus pennsylvanica Marshallgreen ashTreeNative
Grindelia squarrosa (Pursh) Dunalcurlycup gumweedForbNative
Heliopsis helianthoides (L.) Sweetsmooth oxeyeForbNative
Heuchera richardsonii R. Br.Richardson’s alumrootForbNative
Impatiens capensis Meerb.jewelweedForbNative
Lactuca serriola L.prickly lettuceForbIntroduced
Lappula squarrosa (Retz.) Dumort.European stickseedForbIntroduced
Lepidium densiflorum Schrad.common pepperweedForbNative
Liatris aspera Michx.tall blazing starForbNative
Liatris punctata Hook.dotted blazing starForbNative
Linaria vulgaris Mill.butter and eggsForbIntroduced
Linum sulcatum Riddellgrooved flaxForbNative
Lomatium orientale J.M. Coult. & RoseNorthern Idaho biscuitrootForbNative
Lonicera tatarica L.Tatarian honeysuckleShrubIntroduced
Maianthemum racemosum (L.) Linkfeathery false lily of the valleyForbNative
Medicago lupulina L.black medickLegumeIntroduced
Mentha arvensis L.wild mintForbNative
Monarda fistulosa L.wild bergamotForbNative
Muhlenbergia cuspidata (Torr. ex Hook.) Rydb.plains muhlyGraminoidNative
Nuttallanthus canadensis (L.) D.A. SuttonCanada toadflaxForbNative
Oenothera biennis L.common evening primroseForbNative
Oligoneuron rigidum (L.) Smallstiff goldenrodForbNative
Opuntia fragilis (Nutt.) Haw.brittle pricklypearShrubNative
Opuntia humifusa (Raf.) Raf.devil’s-tongueShrubNative
Osmorhiza berteroi DC.sweetcicelyForbNative
Oxalis stricta L.common yellow oxalisForbNative
Pancium capillare L.witchgrassGraminoidNative
Panicum virgatum L.switchgrassGraminoidNative
Parthenocissus tricuspidata (Siebold & Zucc.) Planch.Boston ivyVineIntroduced
Parthenocissus vitacea (Knerr) Hitchc.woodbineVineNative
Phalaris arundinacea L.reed canarygrassGraminoidNative
Physalis virginiana Mill.Virginia groundcherryForbNative
Plantago major L.common plantainForbIntroduced
Plantago rugelii Decne.blackseed plantainForbNative
Poa pratensis L.Kentucky bluegrassGraminoidNative
Polygonatum biflorum (Walter) Elliottsmooth Solomon’s sealForbNative
Polygonum convolvulus L.black bindweedForbIntroduced
Polygonum lapathifolium L.curlytop knotweedForbNative
Polygonum pensylvanicum L.Pennsylvania smartweedForbNative
Polygonum persicaria L.spotted ladysthumbForbIntroduced
Portulaca oleracea L.little hogweedForbIntroduced
Potentilla arguta Purshtall cinquefoilForbNative
Prunus americana MarshallAmerican plumShrubNative
Prunus pumila L.sandcherryShrubNative
Prunus serotina Ehrh.black cherryTreeNative
Prunus virginiana L.chokecherryShrubNative
Quercus macrocarpa Michx.bur oakTreeNative
Ratibida pinnata (Vent.) Barnhartpinnate prairie coneflowerForbNative
Rhus glabra L.smooth sumacShrubNative
Ribes aureum Purshgolden currantShrubNative
Rosa blanda Aitonsmooth roseShrubNative
Rubus occidentalis L.black raspberryVineNative
Rumex crispus L.curly dockForbIntroduced
Schizachyrium scoparium (Michx.) Nashlittle bluestemGraminoidNative
Scirpus atrovirens Willd.green bulrushGraminoidNative
Scutellaria parvula Michx.small skullcapForbNative
Securigera varia (L.) LassencrownvetchLegumeIntroduced
Setaria viridis (L.) P. Beauv.green bristlegrassGraminoidIntroduced
Silene latifolia Poir.bladder campionForbIntroduced
Silphium integrifolium Michx.wholeleaf rosinweedForbNative
Smilax lasioneura Hook.Blue Ridge carrionflowerVineNative
Solanum dulcamara L.climbing nightshadeVineIntroduced
Solidago altissima L.Canada goldenrodForbNative
Sonchus arvensis L.field sowthistleForbIntroduced
Sporobolus compositus (Poir.) Merr.composite dropseedGraminoidNative
Sporobolus heterolepis (A. Gray) A. Grayprairie dropseedGraminoidNative
Sporobolus heterolepis (A. Gray) A. Grayprairie dropseedGraminoidNative
Symphoricarpos occidentalis Hook.western snowberryShrubNative
Symphyotrichum ericoides (L.) G.L. Nesomwhite heath asterForbNative
Symphyotrichum lanceolatum (Willd.) G.L. Nesomwhite panicle asterForbNative
Taraxacum officinale F.H. Wigg.common dandelionForbIntroduced
Thalictrum dasycarpum Fisch. & Avé-Lall.purple meadow-rueForbNative
Toxicodendron rydbergii (Small ex Rydb.) Greenewestern poison ivyVineNative
Tragopogon dubius Scop.yellow salsifyForbIntroduced
Trifolium repens L.white cloverLegumeIntroduced
Ulmus americana L.American elmTreeNative
Urtica dioica L.stinging nettleForbIntroduced
Verbena bracteata Cav. ex Lag. & Rodr.bigbract verbenaForbNative
Verbena hastata L.  swamp verbenaForbNative
Verbena urticifolia L.white vervainForbNative
Viola canadensis L.Canadian white violetForbNative
Vitis riparia Michx.riverbank grapeVineNative
Zanthoxylum americanum Mill.common pricklyashShrubNative

A difficult site to leave, but after lunch at Lange’s Café in town, the group drove south to Blue Mounds.

Inventory Site 10 – Blue Mounds State Park, Rock County, MN

Bison, elk, wolves and prairie chickens roamed this part of Minnesota over 150-200 years ago. Today, a herd of bison resides in the park. The park has a small population of coyotes and a stable deer population. Birdwatchers can catch glimpses of several western species as well as the birds of the tallgrass prairie.http://www.cityofluverne.org/vertical/Sites/%7BFB20A90A-D58D-47BC-8FC7-1ACA9F9D3B27%7D/uploads/%7BAF3E390F-4B2B-4EB8-966C-97C8B91A7F36%7D.JPG

The large Sioux quartzite rock outcrop, first known as “The Mound,” has provided the park area with an exciting past. The cliff appeared blue to settlers going west in the 1860s and 1870s. They named the prominent landmark, the Blue Mound. 

The mystery of the Blue Mound was not restricted to the cliffs.  At the Mound’s southern end is a 1,250-foot long line of rocks aligned in an east-west direction. Who built it and why is unknown. It is known that on the first day of spring and fall, the sunrise and sunset are lined up with this row of stones. 

The team, apparently having a second wind, identified 94 species on the Mound.

Scientific NameCommon NameTypeOrigin
Acer ginnala Maxim.amur mapleTreeIntroduced
Acer saccharinum L.silver mapleTreeNative
Achillea millefolium L.common yarrowForbNative
Agrostis stolonifera L.creeping bentgrassGraminoidIntroduced
Allium stellatum Fraser ex Ker Gawl.autumn onionForbNative
Alyssum alyssoides (L.) L.pale madwortForbIntroduced
Ambrosia psilostachya DC.Cuman ragweedForbNative
Ambrosia trifida L.great ragweedForbNative
Amorpha canescens PurshleadplantLegumeNative
Andropogon gerardii Vitmanbig bluestemGraminoidNative
Apocynum androsaemifolium L.spreading dogbaneForbNative
Arctium minus Bernh.lesser burdockForbIntroduced
Artemisia campestris L.field sagewortForbNative
Artemisia dracunculus L.TarragonForbNative
Artemisia ludoviciana Nutt.white sagebrushForbNative
Ascelpias syriaca L.common milkweedForbNative
Bidens tripartita L.threelobe beggarticksForbNative
Bouteloua curtipendula (Michx.) Torr.sideoats gramaGraminoidNative
Bouteloua gracilis (Willd. ex Kunth) Lag. ex Griffithsblue gramaGraminoidNative
Bromus inermis Leyss.smooth bromeGraminoidIntroduced
Calystegia sepium (L.) R. Br.hedge false bindweedVineNative
Carex L.sedgeGraminoidNative
Cirsium arvense (L.) Scop.Canada thistleForbIntroduced
Cirsium flodmanii (Rydb.) ArthurFlodman’s thistleForbNative
Conyza canadensis (L.) CronquistCanadian horseweedForbNative
Dalea purpurea Vent.purple prairiecloverLegumeNative
Daucus carota L.Queen Anne’s laceForbIntroduced
Dichanthelium oligosanthes (Schult.) GouldScribner’s rosette grassGraminoidNative
Echinacea angustifolia DC.blacksamson echinaceaForbNative
Elymus repens (L.) GouldquackgrassGraminoidIntroduced
Equisetum laevigatum A. Braunsmooth horsestailForbNative
Euthamia graminifolia (L.) Nutt.flat-top goldentopForbNative
Fraxinus pennsylvanica Marshallgreen ashTreeNative
Geum triflorum Purshold man’s whiskersForbNative
Grindelia squarrosa (Pursh) Dunalcurlycup gumweedForbNative
Helianthus maximiliani Schrad.Maximillian sunflowerForbNative
Helianthus petiolaris Nutt.prairie sunflowerForbNative
Hesperostipa spartea (Trin.) BarkworthporcupinegrassGraminoidNative
Heuchera richardsonii R. Br.Richardson’s alumrootForbNative
Juglans nigra L.black walnutTreeNative
Juncus tenuis Willd.poverty rushGraminoidNative
Juniperus virginiana L.eastern redcedarTreeNative
Koeleria macrantha (Ledeb.) Schult.prairie junegrassGraminoidNative
Lactuca serriola L.prickly lettuceForbIntroduced
Lepidium densiflorum Schrad.common pepperweedForbNative
Lespedeza capitata Michx.roundhead lespedezaLegumeNative
Liatris aspera Michx.tall blazing starForbNative
Melilotus officinalis (L.) Lam.yellow sweetcloverLegumeIntroduced
Melilotus officinalis (L.) Lam. (alba)white sweetcloverLegumeIntroduced
Mirabilis hirsuta (Pursh) MacMill.hairy four o’clockForbNative
Morus alba L.white mulberryTreeIntroduced
Oenothera biennis L.common evening primroseForbNative
Oligoneuron rigidum (L.) Smallstiff goldenrodForbNative
Opuntia fragilis (Nutt.) Haw.brittle pricklypearShrubNative
Opuntia humifusa (Raf.) Raf.devil’s-tongueShrubNative
Panicum virgatum L.switchgrassGraminoidNative
Parthenocissus vitacea (Knerr) Hitchc.woodbineVineNative
Pascopyrum smithii (Rydb.) Á. Lövewestern wheatgrassGraminoidNative
Penstemon grandiflorus Nutt.large beardtongueForbNative
Phleum pratense L.timothyGraminoidIntroduced
Plantago major L.common plantainForbIntroduced
Plantago patagonica Jacq.woolly plantainForbNative
Poa compressa L.Canada bluegrassForbIntroduced
Poa pratensis L.Kentucky bluegrassGraminoidNative
Polygonatum biflorum (Walter) Elliottsmooth Solomon’s sealForbNative
Populus deltoides W. Bartram ex Marshall ssp. moniliferaplains cottonwoodTreeNative
Potentilla arguta Purshtall cinquefoilForbNative
Potentilla recta L.sulphur cinquefoilForbIntroduced
Prunus americana MarshallAmerican plumShrubNative
Prunus pumila L.sandcherryShrubNative
Prunus virginiana L.chokecherryShrubNative
Quercus macrocarpa Michx.bur oakTreeNative
Ratibida pinnata (Vent.) Barnhartpinnate prairie coneflowerForbNative
Rhamnus cathartica L.common buckthornShrubIntroduced
Rosa arkansana Porterprairie roseShrubNative
Rubus occidentalis L.black raspberryVineNative
Silphium laciniatum L.compassplantForbNative
Solidago altissima L.Canada goldenrodForbNative
Sonchus arvensis L.field sowthistleForbIntroduced
Sonchus arvensis L.field sowthistleForbIntroduced
Spartina pectinata Bosc ex Linkprairie cordgrassGraminoidNative
Strophostyles helvola (L.) Elliottamberique-beanLegumeNative
Symphoricarpos occidentalis Hook.western snowberryShrubNative
Symphyotrichum ericoides (L.) G.L. Nesomwhite heath asterForbNative
Symphyotrichum lanceolatum (Willd.) G.L. Nesomwhite panicle asterForbNative
Toxicodendron rydbergii (Small ex Rydb.) Greenewestern poison ivyVineNative
Tragopogon dubius Scop.yellow salsifyForbIntroduced
Trifolium pratense L.red cloverLegumeIntroduced
Trifolium repens L.white cloverLegumeIntroduced
Ulmus americana L.American elmTreeNative
Ulmus pumila L.Siberian elmTreeIntroduced
Verbascum thapsus L.common mulleinForbIntroduced
Verbena hastata L.  swamp verbenaForbNative
Verbena urticifolia L.white vervainForbNative
Vitis riparia Michx.riverbank grapeVineNative

The native legume roundhead lespedeza (Lespedeza capitata), found on this site, has multiple uses. The Comanche made a beverage from the leaves. The Dakota, the Omaha, and the Ponca used the stems for a moxa in cases of neuralgia and rheumatism. The small stems were cut in short pieces and attached to the skin by moistening one end with the tongue; the other end was set on fire and allowed to burn down to the skin. The root of roundhead was considered an antidote to poisons that had been swallowed. The exact method of treatment was not specified. 

At one time, this plant was being studied by French scientists for possible medicinal uses. 

Then it was east to Spirit Lake, Iowa, and the town of Okobogi for dinner at Fisherman’s Wharf.  Walleye was the special on the menu.

Day 7 – Saturday, August 24, 1913

We awoke to our first rain of the trek. It continued through mid-morning, limiting our inventory of the Cayler Prairie.  

Fortunately, photographer Oaks was up in time to zip out to the site and capture the sun before it disappeared until mid-afternoon.  

After a quick errand to identify an arboreal hydrangea for the hotel desk clerk, the team drove a short distance west to the next site.

Inventory Site 11 – Cayler Prairie, Dickenson County, IA

Cayler Prairie is a part of the Iowa Prairie Network, a 160-acre native prairie located in Dickinson County near the western edge of the Des Moines Lobe.  The site is used every summer for research by Iowa Lakeside Laboratory students. The prairie has more than 200 species of plants and a variety of animals, including white-tailed jackrabbits, northern harriers, and regal fritillaries. An additional 385 acres of former pasture and cropland adjacent to the prairie were purchased by the Iowa Department of Natural Resources in 1998, with initial help from the Iowa Natural Heritage Foundation.

Cayler is a unique geologic area with knob and kettle topography. The irregular rocky landscape was formed by the glaciers some 12,000 to 14,000 years ago. As the stagnant ice sheet melted, it created a complex of potholes and ridges. Some of the ridges are thought to be eskers, riverbeds that formed inside the melting glacier. As the water melts, it formed channels at weak points and laid down silt and rock that formed long, low hills.

The continuing moderate rain limited our inventory to a windshield survey along the edge of the thigh high grass.  Nevertheless keen (and trained) eyes assisted by binoculars and zoom lenses identified 58 species. 

Scientific NameCommon NameTypeOrigin
Agrostis scabra Willd.rough bentgrassGraminoidNative
Allium stellatum Fraser ex Ker Gawl.autumn onionForbNative
Amaranthus retroflexus L.redroot amaranthForbNative
Ambrosia psilostachya DC.Cuman ragweedForbNative
Ambrosia trifida L.great ragweedForbNative
Amorpha canescens PurshleadplantLegumeNative
Andropogon gerardii Vitmanbig bluestemGraminoidNative
Apocynum androsaemifolium L.spreading dogbaneForbNative
Artemisia ludoviciana Nutt.white sagebrushForbNative
Ascelpias incarnata L.swamp milkweedForbNative
Ascelpias syriaca L.common milkweedForbNative
Asclepias verticillata L.whorled milkweedForb Native
Asparagus officinalis L.garden asparagusForb Introduced
Brickellia eupatorioides (L.) Shinnersfalse bonesetForbNative
Bromus inermis Leyss.smooth bromeGraminoidIntroduced
Calystegia sepium (L.) R. Br.hedge false bindweedVineNative
Cirsium arvense (L.) Scop.Canada thistleForbIntroduced
Cirsium flodmanii (Rydb.) ArthurFlodman’s thistleForbNative
Coreopsis palmata Nutt.stiff tickseedForbNative
Desmodium illinoense A. GrayIllinois ticktrefoilLegumeNative
Echinacea angustifolia DC.blacksamson echinaceaForbNative
Equisetum hyemale L.scouringrush horsetailForbNative
Eupatorium perfoliatum L.common bonesetForbNative
Galium boreale L.northern bedstrawForbNative
Gentiana andrewsii Griseb.closed bottle gentianForbNative
Glycyrrhiza lepidota PurshAmerican licoriceLegumeNative
Helianthus grosseserratus M. Martenssawtooth sunflowerForbNative
Helianthus petiolaris Nutt.prairie sunflowerForbNative
Juncus interior Wiegandinland rushGraminoidNative
Lactuca serriola L.prickly lettuceForbIntroduced
Lactuca tatarica (L.) C.A. Mey.blue lettuceForbNative
Liatris aspera Michx.tall blazing starForbNative
Lotus corniculatus L.bird’s-foot trefoilLegumeIntroduced
Melilotus officinalis (L.) Lam.yellow sweetcloverLegumeIntroduced
Melilotus officinalis (L.) Lam. (alba)white sweetcloverLegumeIntroduced
Monarda fistulosa L.wild bergamotForbNative
Oenothera biennis L.common evening primroseForbNative
Oligoneuron rigidum (L.) Smallstiff goldenrodForbNative
Pancium capillare L.witchgrassGraminoidNative
Panicum virgatum L.switchgrassGraminoidNative
Physalis heterophylla Neesclammy groundcherryForbNative
Prunus americana MarshallAmerican plumShrubNative
Pycnanthemum virginianum (L.) T. Dur. & B.D. Jacks. ex B.L. Rob. & FernaldVirginia mountainmintForbNative
Ratibida pinnata (Vent.) Barnhartpinnate prairie coneflowerForbNative
Rosa arkansana Porterprairie roseShrubNative
Schizachyrium scoparium (Michx.) Nashlittle bluestemGraminoidNative
Setaria viridis (L.) P. Beauv.green bristlegrassGraminoidIntroduced
Solidago altissima L.Canada goldenrodForbNative
Solidago missouriensis Nutt.Missouri goldenrodForbNative
Sorghastrum nutans (L.) NashIndiangrassGraminoidNative
Spartina pectinata Bosc ex Linkprairie cordgrassGraminoidNative
Symphoricarpos occidentalis Hook.western snowberryShrubNative
Symphyotrichum ericoides (L.) G.L. Nesomwhite heath asterForbNative
Taraxacum officinale F.H. Wigg.common dandelionForbIntroduced
Thalictrum dasycarpum Fisch. & Avé-Lall.purple meadow-rueForbNative
Tragopogon dubius Scop.yellow salsifyForbIntroduced
Ulmus pumila L.Siberian elmTreeIntroduced
Verbena hastata L.  swamp verbenaForbNative
Verbena urticifolia L.white vervainForbNative

Part of the rainy morning was spent at the Prairie Chick roadside establishment. One member of the trek commented that he had his all-time best pastry. This Prairie Chick was without a doubt the coffee clache for Iowa’s urban elite on vacation.  The six grass lovers blended right in, well kind of, especially those in their 80s.

From there south through corn, soybeans, smooth brome, and wind turbines to our last inventory.

Inventory Site 12 – Steele Prairie State Preserve, Cherokee County, IA

Erling Jacobson had said there were no chiggers north of the Minnesota – Iowa border.  As a result team members threw caution to the wind and plunged into the grass at Steele Prairie, north of Cherokee, Iowa.

As the inventory wore on and a hint of itching was suspected the four chigger intolerants found their way back to the vehicle, where Oaks photographed the reaction when chigger-laden botanists are spotted on a desolate country road.  Meanwhile Holzworth and Carlson put a wrap on a productive inventory, 74 species.


Scientific NameCommon NameTypeOrigin
Achillea millefolium L.common yarrowForbNative
Allium stellatum Fraser ex Ker Gawl.autumn onionForbNative
Ambrosia psilostachya DC.Cuman ragweedForbNative
Ambrosia trifida L.great ragweedForbNative
Amorpha canescens PurshleadplantLegumeNative
Amorpha fruticosa L.false indigo bushShrubNative
Andropogon gerardii Vitmanbig bluestemGraminoidNative
Anemone cylindrica A. Graycandle anemoneForbNative
Apocynum androsaemifolium L.spreading dogbaneForbNative
Artemisia ludoviciana Nutt.white sagebrushForbNative
Ascelpias incarnata L.swamp milkweedForbNative
Ascelpias syriaca L.common milkweedForbNative
Asparagus officinalis L.garden asparagusForb Introduced
Bromus inermis Leyss.smooth bromeGraminoidIntroduced
Ceanothus americanus L.New Jersey teaShrubNative
Chenopodium pratericola Rydb.desert goosefootForbNative
Cirsium altissimum (L.) Hilltall thistleForbNative
Cirsium arvense (L.) Scop.Canada thistleForbIntroduced
Comandra umbellata (L.) Nutt.bastard toadflaxForbNative
Convolvulus arvensis L.field bindweedForbIntroduced
Coreopsis palmate Nutt.stiff tickseedFortNative
Dalea candida Michx. ex Willd.white prairiecloverLegumeNative
Dalea purpurea Vent.purple prairiecloverLegumeNative
Desmodium illinoense A. GrayIllinois ticktrefoilLegumeNative
Dichanthelium oligosanthes (Schult.) GouldScribner’s rosette grassGraminoidNative
Digitaria sanguinalis (L.) Scop.hairy crabgrassGraminoidIntroduced
Equisetum laevigatum A. Braunsmooth horsestailForbNative
Eriochloa villosa (Thunb.) Kunthhairy cupgrassGraminoidIntroduced
Euphorbia dentata Michx.toothed spurgeForbNative
Fraxinus pennsylvanica Marshallgreen ashTreeNative
Galium boreale L.northern bedstrawForbNative
Geum triflorum Purshold man’s whiskersForbNative
Glycyrrhiza lepidota PurshAmerican licoriceLegumeNative
Helianthus grosseserratus M. Martenssawtooth sunflowerForbNative
Helianthus maximiliani Schrad.Maximillian sunflowerForbNative
Helianthus pauciflorus Nutt. ssp. pauciflorus stff sunflowerForbNative
Heliopsis helianthoides (L.) Sweetsmooth oxeyeForbNative
Hesperostipa spartea (Trin.) BarkworthporcupinegrassGraminoidNative
Heuchera richardsonii R. Br.Richardson’s alumrootForbNative
Lathyrus palustris L. marsh peaLegumeNative
Lespedeza capitata Michx.roundhead lespedezaLegumeNative
Liatris aspera Michx.tall blazing starForbNative
Melilotus officinalis (L.) Lam.yellow sweetcloverLegumeIntroduced
Oenothera biennis L.common evening primroseForbNative
Oligoneuron rigidum (L.) Smallstiff goldenrodForbNative
Oxalis violacea L.violet woodsorrelForbNative
Panicum virgatum L.switchgrassGraminoidNative
Pediomelum argophyllum (Pursh) J. Grimessilverleaf Indian breadrootLegumeNative
Phalaris arundinacea L.reed canarygrassGraminoidNative
Phlox pilosa L.downy phloxForbNative
Physalis virginiana Mill.Virginia groundcherryForbNative
Poa pratensis L.Kentucky bluegrassGraminoidNative
Potentilla arguta Purshtall cinquefoilForbNative
Pycnanthemum virginianum (L.) T. Dur. & B.D. Jacks. ex B.L. Rob. & FernaldVirginia mountainmintForbNative
Ratibida pinnata (Vent.) Barnhartpinnate prairie coneflowerForbNative
Rosa arkansana Porterprairie roseShrubNative
Rudbeckia hirta L.blackeyed SusanForbNative
Rumex altissimus Alph. Woodpale dockForb Native
Rumex crispus L.curly dockForbIntroduced
Sambucus nigra L. ssp. canadensis (L.) R. BolliAmerican black elderberryShrubNative
Schizachyrium scoparium (Michx.) Nashlittle bluestemGraminoidNative
Setaria viridis (L.) P. Beauv.green bristlegrassGraminoidIntroduced
Sorghastrum nutans (L.) NashIndiangrassGraminoidNative
Spartina pectinata Bosc ex Linkprairie cordgrassGraminoidNative
Sporobolus heterolepis (A. Gray) A. Grayprairie dropseedGraminoidNative
Strophostyles helvola (L.) Elliottamberique-beanLegumeNative
Symphyotrichum ericoides (L.) G.L. Nesomwhite heath asterForbNative
Symphyotrichum lanceolatum (Willd.) G.L. Nesomwhite panicle asterForbNative
Symphyotrichum sericeum (Vent.) G.L. Nesomwestern silver asterForbNative
Thalictrum dasycarpum Fisch. & Avé-Lall.purple meadow-rueForbNative
Tradescantia bracteata Smalllongbract spiderwortForbNative
Trifolium pratense L.red cloverLegumeIntroduced
Typha latifolia L.broadleaf cattailForbNative
Verbena urticifolia L.white vervainForbNative

Summary

By early afternoon with the inventory complete, Oaks and the trekkers drove to into Cherokee, the county seat, and enjoyed a pleasant lunch at a downtown family restaurant.  By early evening the group arrived back in Lincoln, the rental car returned, and Bob treated all to a fine eating place close by his home.  After the meal the group parted; the sixth Love of Grass trek was over. 

Why was it so enjoyable? Learning new plants, staying in touch with natural things, being with friends, seeing different landscapes, and recalling the good old days is part of it; these things and more.  The 2014 trek is only six or so months away.  Now that’s something to look forward to.